A Guide to the Study of Fishes, Volume 2 (of 2)Jordan, David Starr
History
A Guide to the Study of Fishes, Volume 2 (of 2)
Jordan, David Starr
Fishes; Fishes -- Bibliography; Zoology -- History
=The Classification of the Bony Fishes.=—The classification of fishes
has been greatly complicated by the variety of names applied to groups
which are substantially but not quite identical one with another. The
difference in these schemes of classification lies in the point of view.
In all cases a single character must be brought to the front; such
characters never stand quite alone, and to lay emphasis on another
character is to make an alteration large or small in the name or in the
boundaries of a class or order. Thus the _Ostariophysi_ with the
_Isospondyli_, _Haplomi_, and a few minor groups make up the great
division of the _Abdominales_. These are fishes in which the ventral
fins are abdominal, that is, inserted backward, so that the pelvis is
free from the clavicle, the two sets of limbs being attached to
different parts of the skeleton. Most of the abdominal fishes are also
soft-rayed fishes, that is, without consecutive spines in the dorsal and
anal fins, and they show a number of other archaic peculiarities. The
Malacopterygians (μαλακός, soft; πτερύξ, fin) of Cuvier therefore
correspond very nearly to the _Abdominales_. But they are not quite the
same, as the spiny-rayed barracudas and mullets have abdominal ventrals,
and many unquestioned thoracic or jugular fishes, as the sea-snails and
brotulids, have lost, through degeneration, all of their fin-spines.
In nearly but not quite all of the Abdominal fishes the slender tube
connecting the air-bladder with the œsophagus persists through life.
This character defines Müller's order of _Physostomi_ (φυσός, bladder;
στόμα, mouth), as opposed to his _Physoclysti_ (φυσός, bladder;
κλεῖστός, closed), in which this tube is present in the embryo or larva
only. Thus the _Thoracices_ and _Jugulares_, or fishes having the
ventrals thoracic or jugular, together correspond almost exactly to the
Acanthopterygians, (ακανθα, spine; πτερύξ, fin), or spiny-rayed fishes
of Cuvier, or to the _Physoclysti_ of Müller. The Malacopterygians, the
_Abdominales_, and the _Physostomi_ are in the same way practically
identical groups. As the spiny-rayed fishes have mostly ctenoid scales,
and the soft-rayed fishes cycloid scales, the _Physostomi_ correspond
roughly to Agassiz's _Cycloidei_, and the _Physoclysti_ to his
_Ctenoidei_.
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