The umbilical vesicle now begins to separate itself more and more from the
abdomen of the foetus, merely a duct of communication passing to that
portion of it which forms the intestinal canal. The first rudiment of the
cord will be found at this separation; its foetal extremity remains for a
long time funnel-shaped, containing, besides a portion of intestine, the
duct of the vesicula umbilicalis, the vasa omphalo-meseraica (the future
vena portæ,) the umbilical vein from the collected venous radicles of the
chorion, and the early trace of the umbilical arteries. These last-named
vessels ramify on a delicate membranous sac of an elongated form which
rises from the inferior or caudal extremity of the embryo, viz. the
_allantois_; whether this is formed by a portion of the mucous layer of
the germinal vesicle, in common with the other abdominal viscera, appears
to be still uncertain: in birds this may be very easily demonstrated as a
vascular vesicle, arising from the extremity of the intestinal canal; and
in mammalia, connected with the bladder by means of a canal called
_urachus_: from its sausage-like shape, it has received the name of
_allantois_.
The existence of an allantois in the human embryo has been long inferred
from the presence of a ligamentous cord extending from the fundus of the
bladder to the umbilicus, like the urachus in animals. But from the
extreme delicacy of the allantois, and from its function ceasing at a very
early period, it had defied all research, until lately when it has been
satisfactorily demonstrated in the human embryo by Baer and Rathke. It
occupies the space between the chorion and amnion, and gives rise
occasionally to a collection of fluid between these membranes, familiarly
known by the name of the liquor amnii spurius, which, strictly speaking is
the liquor allantoidis.
The function of the allantois is still in a great measure unknown. In
animals it evidently acts as a species of receptaculum urinæ during the
latter periods of gestation; but it is very doubtful if this be its use
during the earlier periods. It does not seem directly connected with the
process of nutrition, which at this time is proceeding so rapidly, first
by means of the albuminous contents of the vitellus, or vesicula
umbilicalis, and afterwards by the absorbing radicles of the chorion; but,
from analogy with the structure of the lower classes of animals, it would
appear that it is intended to produce certain changes in the rudimentary
circulation of the embryo, similar to those which, at a later period of
pregnancy, are effected by means of the placenta, and after birth by the
lungs, constituting the great functions of respiration.
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