For it is clear that the growing and developing embryo must obtain, in
some way and from some source, the food-stuff for its nutrition. And
this is effected, among different animals, in one of three ways. Either
the embryo becomes at a very early stage a little, active, voracious,
free-swimming larva, obtaining for itself in these early days of life
its own living; as is the case, for example, with the oyster or the
star-fish. Or the egg from which it is developed contains a large store
of food-yolk, on which it can draw without stint; as is the case with
birds. Or else the embryo becomes attached to the maternal organism in
such a way that it can draw on her for all the nutriment which it may
require; as is the case with the higher mammals.
In both these latter cases the food-material is drawn from the maternal
organism, and is the result of parental sacrifice; but in different
ways. In the case of the bird, the protoplasm of the ovum has acquired
the power of storing up the by-products of its vital activity. The ovum
of such an animal seems at first sight a standing contradiction to the
statement, made some pages back, that the cell cannot grow to any great
extent without undergoing division or fission; and this because volume
tends to outrun surface. For the yolk of a bird's egg is a single cell,
and is often of large size. But when we come to examine carefully these
exceptional cases of very large cells--for what we call the yolk of an
egg is, I repeat, composed of a single cell--we find that the formative
protoplasm is arranged as a thin patch on one side of the yolk in the
case of the bird's egg, or as a thin pellicle surrounding the yolk in
the case of that of the lobster or the insect. All the rest is a product
of protoplasmic life stowed away beneath the patch or within the
pellicle. And this stored material is relatively stable and inert, not
undergoing those vital disruptive changes which are characteristic of
living formative protoplasm. The mass of formative protoplasm, even in
the large eggs of birds, is not very great, and is so arranged as to
offer a relatively extensive surface. All the rest, the main mass of the
visible egg-yolk, is the stored product of a specialized activity of the
formative protoplasm. But all this material is of parental origin--is
elaborated from the nutriment absorbed and digested by the mother.
Public-domain text, read in full here on John Shaqi.
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