Being Well-Born: An Introduction to EugenicsGuyer, Michael F. (Michael Frederic)
Philosophy
Being Well-Born: An Introduction to Eugenics
Guyer, Michael F. (Michael Frederic)
Eugenics; Genetics
=Dominance Not Always Complete.--=As a matter of fact close inspection
shows that in numerous instances dominance is not absolute since traces of
the recessive character may be detectable. For example, in the cross
between smooth and bearded wheat while smoothness is regarded as the
dominant character and beardlessness as the recessive, nevertheless in the
hybrid offspring a slight tendency toward bearding is not infrequently
seen. Or again when horned breeds of cattle are crossed with hornless
ones, a small proportion of such progeny will show traces of imperfect
horns.
In some cases instead of either character dominating the other a form
intermediate between the two parents may result, as we have seen already
in the case of the Andalusian fowl. Thus, certain white-flowered plants
and certain red-flowered plants when crossed produce pink hybrids, and
longheaded and shortheaded wheats when crossed give offspring with heads
of intermediate length. Or again, crosses between white and red cattle may
yield red roans, and between black and white cattle, blue roans.
Thus, while for such pairs of alternative characters as have been studied,
dominance to some considerable degrees at least, seems to be the rule,
still we have gradations down to the intermediate condition, and in some
instances the hybrid with respect to a given character may be unlike
either parent. The things of chief importance in the Mendelian discovery
are the independent, unitary nature of the characters and their
segregation in the offspring of cross-bred forms.
=Modifications of Dominance.--=It should be noted also that there is such
a condition as _delayed dominance_. Davenport found, for example, that
chicks produced by crossing pure white with pure black Leghorn fowls are
speckled black and white, but later in the adult form white becomes
dominant. Likewise conditions of delayed dominance are known in man in
eye-color and notably in color of hair. Some few cases have been recorded
where a character is dominant at one time, recessive at another.
According to Davenport extra toe in fowls may behave in this way.
=Mendel's Own Work.--=Mendel[2] himself worked out his principles on seven
pairs of characters which he found in common culinary peas. Placing the
dominant characters first, these may be enumerated as follows: (1) Tall by
dwarf; (2) green pod (unripe) by yellow; (3) pod inflated by pod
constricted between the individual peas; (4) flowers arranged along the
axis of the plant by flowers bunched together at the top; (5) seed skin
colored by seed skin white; (6) cotyledons yellow by cotyledons green; (7)
seed rounded by seed wrinkled.
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