But, quite independently of this peculiar growth habit, the stems and
often whole plants do show response to light and many times the
response, in its effects, cannot be distinguished from geotropism.
Perhaps the most homely illustration of this is the common house
geranium which, no matter how often it is turned, always grows toward
the window, and if not turned at all becomes hopelessly lopsided, with
the leaves all bending sharply toward the light. Trees growing on a
cliffside, while always growing upward, nearly always may be seen
bending away from the cliff where light is scarce and toward the
unobstructed light. The position of hundreds of twigs and branches on
any tree have been dictated by their exposure to light, and the habit of
practically all trees in the forest of being clear of branches for many
feet from the ground is another illustration of the profound effect of
light. In the latter case the taller the trees the farther from the
ground are the first branches, and in the big trees of California the
first branches are frequently over a hundred feet from the ground. In
their young stages all these trees were furnished with branches, the
leaves of which in their day performed their appointed tasks. But in the
strife and hurry of the crowns of the forest to overreach their
neighbors these lower branches, from the bottom upward, gradually die
off. So inexorable is the plant’s demand for light, that these lower
branches, in spite of being nearest the source of their food from the
roots, are doomed to be killed. Nature plays no favorites and these
lower branches, once the pride and support of the young tree, are
ruthlessly dropped off when they can no longer play the game. This
wholesale slaughter of lower branches in a forest, more complete than
any pruning by man could ever be, gives us, if the story of the factory
leaf has not already done so, some conception of the part played by
light in the plant world.
Public-domain text, read in full here on John Shaqi.
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