For example, the fossil mammals from the Australian
caves are nearly allied to the modern kangaroos, phalangers, and
wombats; and the gigantic extinct sloths and armadillos of South America
are reproduced in their smaller representatives at the present day. So,
too, the moa of New Zealand was a huge apteryx; and the birds
disentombed from the bone-caves of Brazil show close affinities to the
toucans and jacanars that still scream and flit in countless flocks
among Brazilian forests. The obvious implication is that the animals now
inhabiting any given area are the modified descendants of those that
formerly inhabited it. 'On the theory of descent with modification, the
great law of the succession of the same types within the same areas is
at once explained.'
This last consideration leads us up to the argument from Geographical
Distribution. In considering the various local faunas and floras on the
face of the globe, no point strikes one more forcibly than the fact that
neither their similarities nor their dissimilarities can be accounted
for by climate or physical conditions. The animals of South Africa do
not in the least resemble the animals of the corresponding belt of South
America; the Australian beasts and birds and trees are utterly unlike
those of France and Germany; the fishes and crustaceans of the Pacific
at Panama are widely different from those of the Caribbean at the same
point, separated from them only by the narrow belt of intervening
isthmus. On the other hand, within the same continuous areas of sea or
land, however great the differences of physical conditions, we find
everywhere closely related types in possession of the most distinct and
varied situations. On the burning plains of La Plata we get the agouti
and the bizcacha as the chief rodents; we ascend the Cordillera, and
close to the eternal snows we discover, not hares and rabbits like those
of Europe, but a specialised chilly mountain form of the same distinctly
South American type. We turn to the rivers, and we see no musk-rat or
beaver, but the coypu and capybara, slightly altered varieties of the
original bizcacha ancestor. Australia has no wolf, but it has instead
fierce and active carnivorous marsupials; it has no mice, but some of
its tiny kangaroo-like creatures fulfil analogous functions in its
animal economy. Everywhere the evidence points to the conclusion that
local species have been locally evolved from pre-existing similar
species. The oceanic isles, of which Darwin had had so large an
experience, and especially his old friends the Galapagos, come in
usefully for this stage of the question. They are invariably inhabited,
as Darwin pointed out, and as Wallace has since abundantly shown in the
minutest detail, by waifs and strays from neighbouring continents,
altered and specialised by natural selection in accordance with the
conditions of their new habitat. As a rule, they point back to the
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