Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
The question to be settled is whether acquired characters are ever
transmitted by heredity. Now suppose, for the sake of argument, that
acquired characters are transmitted by heredity--though not so fully and
not so certainly as congenital characters--how is this fact to be proved
to the satisfaction of Weismann and his followers? First of all they
answer,--Assuredly by adducing experimental proof of the inheritance of
injuries, or mutilations. But in making this answer they appear to
forget that Darwin has already shown its inefficiency. That the
self-styled Neo-Lamarckians have been much more unguarded in this
respect, I fully admit; but it is obviously unfair to identify Darwin's
views with those of a small section of evolutionists, who are really as
much opposed to Darwin's teaching on one side as is the school of
Weismann on the other. Yet, on reading the essays of Weismann
himself--and still more those of his followers--one would almost be led
to gather that it is claimed by him to have enunciated the distinction
between congenital and acquired characters in respect of
transmissibility; and therefore also to have first raised the objection
which lies against the theory of Pangenesis in respect of the
non-transmissibility of mutilations. In point of fact, however, Darwin
is as clear and decided on these points as Weismann. And his answer to
the obvious difficulty touching the non-transmissibility of mutilations
is, to quote his own words, "the long-continued inheritance of a part
which has been removed during many generations is no real anomaly, for
gemmules formerly derived from the part are multiplied and transmitted
from generation to generation[29]." Therefore, so far as Darwin's theory
is concerned, the challenge to produce evidence of the transmission of
injuries is irrelevant: it is no more a part of Darwin's theory than it
is of Weismann's to maintain that injuries _are_ transmitted.
[29] _Variation under Domestication_, ii. 392.
Public-domain text, read in full here on John Shaqi.
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