=145. Chlorophyll is a pigment which resides in the chloroplast.=—That
the chlorophyll is a coloring substance which resides in the
chloroplastid, and does not form the body itself, can be demonstrated
by dissolving out the chlorophyll when the framework of the
chloroplastid is apparent. The green parts of plants which have been
placed for some time in alcohol lose their green color. The alcohol
at the same time becomes tinged with green. In sectioning such plant
tissue we find that the chlorophyll bodies, or chloroplastids as they
are more properly called, are still intact, though the green color is
absent. From this we know that chlorophyll is a substance distinct from
that of the chloroplastid.
=146. Chlorophyll absorbs energy from sunlight for photosynthesis.=—It
has been found by analysis with the spectroscope that chlorophyll
absorbs certain of the rays of the sunlight. The energy which is thus
obtained from the sun, called _kinetic_ energy, acts on the molecules
of CH₂O₃, separating them into molecules of C, H, and O. (When the
CO₂ from the air enters the plant cell it immediately unites with
some of the water, forming carbonic acid = CH₂O₃.) After a series of
complicated chemical changes starch is formed by the union of carbon,
oxygen, and hydrogen. In this process of the reduction of the CH₂O₃ and
the formation of starch there is a surplus of oxygen, which accounts
for the giving off of oxygen during the process.
=147. Rays of light concerned in photosynthesis.=—If a solution
of chlorophyll be made, and light be passed through it, and this
light be examined with the spectroscope, there appear what are called
absorption bands. These are dark bands which lie across certain
portions of the spectrum. These bands lie in the red, orange, yellow,
green, blue, and violet, but the bands are stronger in the red, which
shows that chlorophyll absorbs more of the red rays of light than of
the other rays. These are the rays of low refrangibility. The kinetic
energy derived by the absorption of these rays of light is transformed
into potential energy. That is, the molecule of CH₂O₃ is broken up, and
then by a different combination of certain elements starch is formed.[8]
[8] In the formation of starch during photosynthesis the separated
molecules from the carbon dioxide and water unite in such a way that
carbon, hydrogen, and oxygen are united into a molecule of starch. This
result is usually represented by the following equation: CO₂ + H₂O =
CH₂O + O₂. Then by polymerization 6(CH₂O) = C₆H₁₂O₆ = grape sugar.
Then C₆H₁₂O₆-H₂O = C₆H₁₀O₅ = starch. It is believed, however, that the
process is much more complicated than this, that several different
compounds are formed before starch finally appears, and that the
formula for starch is much higher numerically than is represented by
C₆H₁₀O₅.
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