Elements of Structural and Systematic Botany: For High Schools and Elementary College Courses — John Shaqi
Elements of Structural and Systematic Botany: For High Schools and Elementary College CoursesCampbell, Douglas Houghton
Science
Elements of Structural and Systematic Botany: For High Schools and Elementary College Courses
Campbell, Douglas Houghton
Botany -- Textbooks
The leaves (Fig. 52, _A_, _B_) are rounded in outline unequally,
two-lobed, and arranged in two rows on the upper side of the stem, so
closely overlapping as to conceal it entirely. On the under side are
similar but smaller leaves, less regularly disposed. The stems branch
at intervals, the branches spreading out laterally so that the whole
plant is decidedly flattened. On the under side are fine, whitish
hairs, that fasten it to the substratum. If we examine a number of
specimens, especially early in the spring, a difference will be
observed in the plants. Some of them will be found to bear peculiar
structures (Fig. 52, _C_, _D_), in which the spores are produced.
These are called "sporogonia." They are at first globular, but when
ripe open by means of four valves, and discharge a greenish brown mass
of spores. An examination of the younger parts of the same plants will
probably show small buds (Fig. 54, _H_), which contain the female
reproductive organs, from which the sporogonia arise.
[Illustration: FIG. 52.--_A_, part of a plant of a leafy liverwort
(_Madotheca_), × 2. _B_, part of the same, seen from below, × 4. _C_,
a branch with two open sporogonia (_sp._), × 4. _D_, a single
sporogonium, × 8.]
On other plants may be found numerous short side branches (Fig. 53,
_B_), with very closely set leaves. If these are carefully separated,
the antheridia can just be seen as minute whitish globules, barely
visible to the naked eye. Plants that, like this one, have the male
and female reproductive organs on distinct plants, are said to be
"diœcious."
A microscopical examination of the stem and leaves shows their
structure to be very simple. The former is cylindrical, and composed
of nearly uniform elongated cells, with straight cross-walls. The
leaves consist of a single layer of small, roundish cells, which,
like those of the stem, contain numerous rounded chloroplasts, to
which is due their dark green color.
The tissues are developed from a single apical cell, but it is
difficult to obtain good sections through it.
The antheridia are borne singly at the bases of the leaves on the
special branches already described (Fig. 53, _A_, _an._). By
carefully dissecting with needles such a branch in a drop of water,
some of the antheridia will usually be detached uninjured, and may
be readily studied, the full-grown ones being just large enough to
be seen with the naked eye. They are globular bodies, attached by a
stalk composed of two rows of cells. The globular portion consists
of a wall of chlorophyll-bearing cells, composed of two layers
below, but single above (Fig. 53, _C_). Within is a mass of
excessively small cells, each of which contains a spermatozoid. In
the young antheridium (_A_, _an._) the wall is single throughout,
and the central cells few in number. To study them in their natural
position, thin longitudinal sections of the antheridial branch
should be made.
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