Evolution and Classification of the Pocket Gophers of the Subfamily GeomyinaeRussell, Robert J.
Science
Evolution and Classification of the Pocket Gophers of the Subfamily Geomyinae
Russell, Robert J.
Pocket gophers
_Diagnosis._--Small Geomyinae; lacking specializations of more
advanced tribes; upper incisors smooth, at least in _Pliosaccomys_;
molariform teeth always rooted and having crowns of medium height;
enamel investment of cheek teeth complete and uninterrupted in all
stages of wear; crowns of molars primitively biprismatic, having two
columns united at mid-points, thus forming narrow isthmus separating
lateral re-entrant folds as in premolars, and, with wear, also uniting
secondarily at protomeres (with exception of third lower molars),
consequently, isolating remnant of that inflection as shallow fossette
(columns uniting first at protomeres in _Pliosaccomys_); anterior and
posterior columns of first and second molars, both above and below,
becoming progressively united into one column in advanced Dikkomyini
(early and middle Pliocene), but m3 (M3 unknown) retaining primitive
biprismatic pattern, with columns joined at centers but never at
protomeres (for details of dentition see generic accounts); mandible
stout, its angle mostly above mandibular tooth-row; masseteric ridge
low; basitemporal fossa barely discernable in some fragments of
_Pliosaccomys_; postcranial skeleton unknown.
Key to the Genera of the Tribe Dikkomyini
A Molars biprismatic throughout life; anterior and posterior
lophs of first and second molars in pre-final stages of wear
uniting first at their mid-points and later at edge of
protomeres; anterior lophid of lower premolar having distinct
anteroexternal inflection. Genus _Dikkomys_ p. 516
A' First and second molars becoming monoprismatic in final
(adult?) stages of wear, biprismatic only in pre-final stages
of wear; third molars persistently biprismatic; anterior and
posterior lophs of first and second molars uniting first at
edge of protomeres; anterior lophid of lower premolar lacking
anteroexternal inflection. Genus _Pliosaccomys_ p. 517
Genus =Dikkomys= Wood
1936. _Dikkomys_ Wood, Amer. Mus. Novit., 866:26, July 2.
_Type._--_Dikkomys matthewi_ Wood, 1936, from Lower Harrison deposits
near Agate, Sioux County, Nebraska.
_Chronologic range._--Early Miocene, from early Arikareean (Lower
Harrison local fauna of Nebraska) to middle Miocene, late
Hemingfordian (Upper Rosebud local fauna, South Dakota, and the Deep
River Formation, Montana). According to MacDonald (1963:149-150), the
Upper Rosebud is middle Miocene rather than early Miocene.
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