On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan regionWallace, Alfred Russel
Science
On the phenomena of variation and geographical distribution as illustrated by the Papilionidæ of the Malayan region
Wallace, Alfred Russel
Papilionidae -- Malaysia -- Geographical distribution; Papilionidae -- Malaysia -- Variation
│ │ │ ║ Seventy-two, Austro-Malayan Region. │
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The exceeding richness of the Malayan region in these fine insects is
seen by comparing the number of species found in the different tropical
regions of the earth. From all Africa only 33 species of _Papilio_ are
known; but as several are still undescribed in collections, we may raise
their number to about 40. In all tropical Asia there are at present
described only 65 species, and I have seen in collections but two or
three which have not yet been named. In South America, south of Panama,
there are 120 species, or about the same number as I make in the Malayan
region; but the area of the two countries is very different; for while
South America (even excluding Patagonia) contains 5,000,000 square
miles, a line encircling the whole of the Malayan islands would only
include an area of 2,700,000 square miles, of which the land-area would
be about 1,000,000 square miles. This superior richness is partly real
and partly apparent. The breaking up of a district into small isolated
portions, as in an archipelago, seems highly favourable to the
segregation and perpetuation of local peculiarities in certain groups;
so that a species which on a continent might have a wide range, and
whose local forms, if any, would be so connected together that it would
be impossible to separate them, may become by isolation reduced to a
number of such clearly defined and constant forms that we are obliged to
count them as species. From this point of view, therefore, the superior
number of Malayan species may be considered as apparent only. Its true
superiority is shown, on the other hand, by the possession of three
genera and twenty groups of Papilionidæ against a single genus and eight
groups in South America, and also by the much greater average size of
the Malayan species. In most other families, however, the reverse is the
case, the South American _Nymphalidæ_, _Satyridæ_, and _Erycinidæ_ far
surpassing those of the East in number, variety, and beauty.
The following list, exhibiting the range and distribution of each group,
will enable us to study more easily their internal and external
relations.
_Range of the Groups of Malayan_ Papilionidæ.
Ornithoptera.
1. _Priamus_-group. Moluccas to Woodlark Island.
2. _Pompeus_-group. Himalayas to New Guinea (Celebes, maximum).
3. _Brookeana_-group. Sumatra and Borneo.
Papilio.
4. _Nox_-group. North India, Java, and Philippines.
5. _Coon_-group. North India to Java.
6. _Polydorus_-group. India to New Guinea and Pacific.
7. _Ulysses_-group. Celebes to New Caledonia.
8. _Peranthus_-group. India to Timor and Moluccas (India, max.).
9. _Memnon_-group. India to Timor and Moluccas (Java, max.).
10. _Helenus_-group. Africa and India to New Guinea.
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