Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
Whilst this dipleuric or bilateral (commonly obliquely ovate or nearly
triangular) form of the sagittal ring is distinctly preserved in by far the
greater number of #Stephoidea#, it is replaced in some few genera by a more
regular, amphithect, diphragmatic, or biradial form. In this case we may
often suppose a primary regularity to exist, the dorsal and ventral parts
being not yet differentiated, as in _Archicircus_ and _Lithocircus_,
_Zygostephanus_ and _Protympanium_, and perhaps also in some other forms.
But in other cases the regularity is, on the contrary, secondary, being
derived from original bilateral forms.
The rod of the sagittal ring is either cylindrical (with circular
transverse section) or angular (commonly with triangular transverse
section). In nearly all #Stephoidea# (with very few exceptions) branches or
apophyses are developed from the ring, regularly disposed and often of
great morphological importance. Commonly these {934}apophyses are developed
in pairs, growing symmetrically on both sides of the ring. The most
important of these apophyses are:--(1) Basal apophyses, arising from the
basal hole of the ring; (2) mitral apophyses, arising from the apical pole;
(3) dorsal apophyses, arising from the middle of the dorsal rod; and (4)
ventral apophyses, arising from the middle of the ventral rod. The two
former arise in the principal axis, the two latter in the sagittal axis of
the body. Very frequently the latter pair is replaced by two pairs of
transverse branches, one inferior (mandibular) and one superior (orbital).
The apophyses of the ring are either simple or branched, often very large,
richly ramified, and give origin to a number of further products.
Whilst in the Stephanida the primary sagittal ring alone represents the
whole skeleton, it produces in all other #Stephoidea# one or more secondary
rings. The most important of these is the horizontal basal ring, appearing
first in the Semantida (Pl. 92). From the base of the sagittal ring there
arise in the horizontal basal plane two pairs of lateral branches or "basal
apophyses." The curved opposite branches of the corresponding pairs become
united on each side of the primary ring (right and left), and so produce a
second, horizontal ring, perpendicular to the former. This basal ring
encloses two paired basal gates, which are enclosed on the medial side by
the basal rod of the sagittal ring, and on the lateral side by two united
apophyses (_Semantis_, Pl. 92, figs. 1, 2). These two primary basal gates
are of the greatest morphological importance; we call them the "jugular
gates or jugular pores" (in the description of Bütschli, the pores I, _loc.
cit._, p. 498). The dorsal pair of basal apophyses (on their posterior
edge) are the coracal rods, _e_ (rods _e_ of Bütschli); the opposite
ventral pair (on their anterior edge) are the clavicular or furcular rods,
_f_ (rods _e_1 of Bütschli); compare Pls. 92-95, and their explanation.
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