Another possibility is that the birds, which are less differentiated
from their ancestral stocks, may be less differentiated because of
suppression of newly evolved characters by dilutions, which result from
interbreeding with new birds, which may be arriving at irregular
intervals from the ancestral home. Interbreeding of the resident
population with newcomers may overshadow any modifications which might
have appeared as a result of insular isolation, especially modifications
which have little adaptive significance. One would suspect, from their
modifications, that _Rallus owstoni_, _Metabolus rugensis_, _Corvus
kubaryi_, and other endemic forms have experienced less of this
"dilution," than such birds as _Rallus philippensis pelewensis_,
_Artamus leucorhynchus pelewensis_, _Myzomela cardinalis_, and others.
Murphy (1938) mentions this "dilution" effect in his discussion of
"strong" and "weak" subspecies among warblers of the Marquesas. He
writes that "strong" subspecies may develop if the birds are present on
islands which are upwind from islands containing related subspecies. The
wind acts to block interisland migration in these weak-flyers. On the
other hand, "weak" subspecies may show the effect of "dilution," being
situated on islands downwind from islands containing related
subspecies. The direction of the wind acts to aid the weak flyers to
move to the downwind islands and continually "dilute" the resident
subspecies. Similar examples can be cited for Micronesian birds. Hesse,
Allee, and Schmidt (1937:87) write, "Endemism on islands is most
frequent in forms for which the difficulty of reaching the island is
most extreme, so that new increments of the parent form are unlikely to
follow."
Employing the criteria mentioned above, the birds of Micronesia can be
tentatively divided into four groups as regards the relative time when
they arrived at the islands:
1. Birds of ancient colonizations which reached certain individual
islands, became modified, and dispersed no farther. Examples are
_Aphanolimnas_, _Rallus owstoni_, _Aplonis corvinus_, _Metabolus
rugensis_, and _Corvus kubaryi_.
2. Birds of ancient colonizations which reached or dispersed through a
number of islands but are now restricted to relatively few islands.
Examples are _Ducula oceanica_, _Ptilinopus porphyraceus_, _Megapodius
lapérouse_, _Asio flammeus_, and _Acrocephalus luscinia_.
3. Birds of ancient, or possibly more recent, colonizations which
initially reached or subsequently dispersed to many of the islands of
Micronesia possessing habitat suitable for them. Examples are _Myzomela
cardinalis_, the two species of _Halcyon_, _Aplonis opacus_, and
_Zosterops conspicillata_.
4. Birds of rather recent colonizations, which may have reached only a
few islands and are relatively unmodified from their parental stocks.
Examples are _Artamus leucorhynchus_, _Caprimulgus indicus_,
_Poliolimnas cinereus_, and _Nycticorax caledonicus_.
FACTORS CAUSING DISPERSAL
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account