The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
This question is, however, wrapped up in a wider one. Its solution depends
upon the ancestry of mammals. If the Mammalia are to be derived from
reptiles with simple conical teeth, then the first stage in the development
of trituberculy is proved. On the other hand, however, the evidence is
gradually growing that the Theromorpha represent more nearly than any
non-mammalian group with which we are acquainted the probable ancestral
form of the mammals. These animals offer some support to both the leading
views. _Cynognathus_ had triconodont teeth which, as will be pointed out
later, are a theoretically intermediate stage in the evolution of
tritubercular teeth; on the other hand, the teeth of _Diademodon_ and some
others are multituberculate, and have been very properly compared to the
multitubercular teeth of such primitive mammalia as the Ornithorhynchus.
Professor Osborn is no doubt correct in italicising a remark of an
anonymous writer in _Science_ to the effect that in _Diademodon_ the teeth,
though multitubercular, show the prevalence of three cusps arranged in the
tritubercular fashion. {57} But this may be only a proof that the
multitubercular antedates the tritubercular. It may be, indeed, that the
mammalian tooth was already differentiated among the mammal-like Saurians
and that from such a form as _Cynognathus_ the Eutheria and other forms in
which a tritubercular arrangement can be detected were evolved, and from
such form as _Tritylodon_ the Monotrematous branch of the mammals. This way
of looking at the matter harmonises a much-disputed question, but involves
a diphyletic origin of the mammals--an origin which for other reasons is
not without its supporters.
We shall now attempt to give a general idea of the facts and arguments
which support or tend to support "trituberculy." As a matter of fact the
name is inaccurate; for the holders of this view do not derive the
mammalian molar from a trituberculate condition, but in the first place
from a simple cone such as that of a crocodile!
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