The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
(_n_) The _pectoral girdle_ (187) is a very simple and uniform structure,
although the details of the association of the elements still remain to
be determined. A single, median, usually large and elongate interclavicle
occupies the ventral line of the chest. This is morphologically the same
element which occurs in the middle line of the chest of the lizards.
It is a dermal bone and is usually, especially among the Microsauria
(462), highly sculptured. It varies considerably in size and shape, but
is remarkably uniform throughout the various groups. Lying anterior to
the interclavicle and overlapping its antero-lateral margins lie the two
clavicles, which are usually diamond-shaped and are sculptured, dermal
bones. The position of the coracoid is still uncertain, and in fact its
clear association in the pectoral girdle of these species is still a
question. It seems to be constant in the European (186, 251) species and
is usually represented by a small rounded plate of bone, which in life no
doubt had a large amount of cartilage to form its borders. A cleithrum
(285) has been ascribed to one of the Linton, Ohio, species (plate 15,
fig. 3) by Jaekel (347), but this needs confirmation. An osseous scapula
is usually present, resembling the scapula of modern salamanders, in that
it was largely embedded in cartilage. The position of the pectoral girdle
is largely a matter of doubt, especially for the American species. After
death and before fossilization the girdle was always moved by post-mortem
shifting, so that its exact relation to the ribs and vertebral column
is still in doubt. Credner (186) has restored the pectoral girdle close
behind the head, which would cause an amount of rigidity in the body which
probably did not exist.
(_o_) The _arm_ consists of the humerus, radius, ulna, and 4 digits.
The characters of the arm-bones are such as is constant among primitive
animals and developing mammals. The osseous portion is perichondral.
Epiphyses are totally lacking and it is doubtful if the endochondrium was
at all ossified. The digits are often terminated by ungual phalanges,
although usually the terminal phalanx was merely embedded in the web of
the foot; and among the terrestrial forms a claw was well developed. An
osseous carpus is not known in the species from the Coal Measures. Its
impression indicates a broad hand, well adapted for swimming.
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