The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
The gastric mill begins in the hinder half of the cardiac division.
Here, on the upper wall of the stomach, we see a broad transverse
calcified bar (figs. 9–11, _c_) from the middle of the hinder part of
which another bar (_uc_), united to the first by a flexible portion,
is continued backwards in the middle line. The whole has, therefore,
somewhat the shape of a cross-bow. Behind the first-mentioned piece,
the dorsal wall of the stomach is folded in, in such a manner as
to give rise to a kind of pouch; and the second piece, or what we
may call the handle of the crossbow, lies in the front wall of this
pouch. The end of this piece is dense and hard, and its free surface,
which looks into the top of the cardiac chamber, is raised into two
oval, flattened convex surfaces (_t_). Connected by a transverse
joint with the end of the handle of the crossbow, there is another
solid bar, which ascends obliquely forwards in the back wall of the
pouch (_pp_). The end which is articulated with the handle of the
crossbow is produced into a strong reddish conical tooth (_mt_),
curved forwards and bifurcated at the summit; consequently, when the
cavity of the stomach is inspected from the fore part of the cardiac
pouch (fig. 9, B), the two-pointed curved tooth (_mt_) is seen
projecting behind the convex surfaces (_t_), in the middle line, into
the interior of that cavity. The joint which connects the handle of
the crossbow with the hinder middle piece is elastic; hence, if the
two are straightened out, they return to their bent disposition as
soon as they are released. The upper end of {55} the hinder middle
piece (_pp_) is connected with a second flat transverse plate which
lies in the dorsal wall of the pyloric chamber (_p_). The whole
arrangement, thus far, may be therefore compared to a large cross-bow
and a small one, with the ends of their handles fastened together by
a spring joint, in such a manner that the handle of the one makes an
acute angle with the handle of the other; while the middle of each
bow is united with the middle of the other by the bent arm formed
by the two handles. But, in addition to this, the outer ends of the
two bows are also connected together. A small, curved, calcified bar
(_pc_) passes from the outer end of the front crosspiece downwards
and outwards in the wall of the stomach, and its hinder and lower
extremity is articulated with another larger bar (_zc_) which runs
upwards and backwards to the hinder or pyloric crosspiece, with
which it articulates. Internally, this piece projects into the
cardiac cavity of the stomach as a stout elongated reddish elevation
(_lt_), the surface of which is produced into a row of strong sharp,
transverse ridges, which diminish in size from before backwards, and
constitute a crushing surface almost like that of the grinder of an
elephant. Thus, when the front part of the cardiac cavity is cut
away, not only are the median teeth already mentioned seen, but, on
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