The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
The vegetation appertaining to the Sporangites has not been distinctly
recognised. I have, however, found in one of the Brazilian specimens two
sporocarps attached to what seems a fragment of a cellular frond, and
numerous specimens of the supposed Algæ, named _Spirophyton_, are found
in the shales, but there is no evidence of any connection of this plant
with the _Protosalvinia_.
Modern Rhizocarps present considerable differences as to their vegetative
parts. Some, like _Pilularia_, have simple linear leaves; others, like
_Marsilea_, have leaves in whorls, and cuneate in form; while others,
like _Azolla_ and _Salvinia_, have frondose leaves, more or less pinnate
in their arrangement. If we inquire as to fossils representing these
forms of vegetation, we shall find that some of the plants to be noticed
in the immediate sequel may have been nearly allied to the Rhizocarps.
In the mean time I may state that I have proposed the generic name
_Protosalvinia_ for these curious macrospores and their coverings, and
have described in the paper in the "Bulletin of the Chicago Academy of
Sciences," already quoted, five species which may be referred to this
genus.
These facts lead to inquiries as to the origin of the bituminous matter
which naturally escapes from the rocks of the earth as petroleum and
inflammable gas, or which may be obtained from certain shales in
these forms by distillation. These products are compounds of carbon
and hydrogen, and may be procured from recent vegetable substances by
destructive distillation. Some vegetable matters, also, are much richer
in carbon and hydrogen than others, and it is a remarkable fact that
the spores of certain cryptogamous plants are of this kind, as we see in
the inflammable character of the dry spores of Lycopodium; and we know
that the slow putrefaction of such material underground effects chemical
changes by which bituminous matter can be produced. There is, therefore,
nothing unreasonable in the supposition advanced by Prof. Orton, that the
spores so abundantly contained in the Ohio black shales are important
or principal sources of the bituminous matter which they contain.
Microscopic sections of this shale show that much of its material
consists of the rich bituminous matter of these spores (Fig. 16). At the
same time, while we may trace the bitumen of these shales, and of some
beds of coal, to this cause, we must bear in mind that there are other
kinds of bituminous rocks which show no such structures, and may have
derived their combustible material from other kinds of vegetable matter,
whether of marine or of land plants. We shall better understand this when
we have considered the origin of coal.
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