The test, then, of a mutation is not so much the amount of deviation as
the degree in which it is inherited. Mutations show no tendency to a
gradual return to the mean of the parent species; fluctuating variations
do display such a tendency. A mutation consists, as M. E. East says, in
the production of a new mode or centre for linear fluctuation; it is, as
it were, a shifting of the centre of gravity; the centre about which
those fluctuations which we call continuous variations occur.
As it is of considerable importance thoroughly to grasp the true nature
of mutations or discontinuous variations, and as some writers do not
appear to realise wherein lies the essential difference between the two
kinds of variation, we will, at the risk of appearing tedious, give a
further illustration. Let A be a species of bird of which the average
length of the wing is 20 inches, and let us suppose that individuals
belonging to that species occur in which the length of the wing varies as
much as 3 inches each side of the mean; thus it is possible to find
individuals of this species with a wing as short as 17 inches, or as long
as 23 inches. Let B be another species of which the average length of the
wing is 17 inches, and let us suppose that a 3-inch variation on each
side of the mean be found to occur. Individuals belonging to species B
will occur which have a wing as short as 14 inches, or as long as 20
inches. Thus some individuals of the short-winged species will have
longer wings than certain individuals of the long-winged species.
Similarly, certain individuals of a species which display a mutation may
show less deviation from the mean than some individuals showing a very
pronounced fluctuating variation. In other words, even as by measuring
the length of wing in the above example it was not always possible to say
whether a given individual belonged to species A or B, so is it not
always possible to say by looking at an individual that shows a
considerable departure from the mean whether that departure is due to a
mutation or a fluctuating variation.
Law of Regression
It is only by watching the effect of the peculiarity on the offspring of
its possessor that we are able to determine the nature of the variation.
Where the peculiarity is due to a fluctuating variation the offspring
will display the peculiarity in a diminished degree; but if the
peculiarity be due to a mutation, the offspring are likely to display it
in as marked a degree as the parent.
Fritz Müller and Galton conducted independently enquiries into the amount
of the regression shown by the progeny of parents which have deviated
from the average by fluctuating variation.
Müller experimented with Indian corn; Galton with the sweet pea.
Public-domain text, read in full here on John Shaqi.
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