The physiology of digestion considered with relation to the principles of dieteticsCombe, Andrew
Science
The physiology of digestion considered with relation to the principles of dietetics
Combe, Andrew
Diet; Digestion
In ruminating animals, such as the sheep and ox, the stomach, as will
be seen from the annexed figure, not only is large, to adapt it to the
bulky nature of their food, but is complicated in its structure, to fit
it for effecting the great changes which vegetable aliment requires to
undergo before it can be converted into blood. It may indeed be said
to consist of four distinct stomachs conjoined. In the first of these,
AA, termed the _paunch_, the herbage is deposited when first
swallowed after hasty and ineffectual mastication. It there undergoes
a kind of maceration or steeping in a fluid provided for the purpose;
after which it passes from the paunch into a smaller bag, called the
_reticulum_, or bonnet, B, which, in some animals, such as the
camel and dromedary, is designed exclusively as a reservoir for water,
which being there stored up in large quantities, ready for use when
wanted, fits them in a wonderful manner for travelling through the
arid deserts where no water is to be obtained, and where, without some
such provision, they would of course soon perish. So admirably is the
reticulum adapted for this special purpose, that the water contained
in it undergoes little or no change either in quality or quantity,
although if it were collected in the ordinary digesting stomach, it
would be entirely absorbed in the course of a few minutes. It is not
even mixed with the food which is swallowed after it, as the animal has
the power of directing solids at once into the other cavities. From the
reticulum the alimentary mass is again returned to the mouth, there
to be thoroughly masticated and mixed with the saliva; after which it
descends a second time through the gullet: but instead of passing,
as before, into the paunch, it enters the third bag, _omasum_,
or many-plies, C, where it undergoes farther changes, and is then
transmitted to the fourth portion D, adjoining the pylorus, and named
_ab-omasum_, or red-bag. The last portion is exactly similar in
structure and in function to the simple stomach of man and the other
mammalia, and is in fact the true stomach, the other three being merely
preparatory organs.
The first part of the process, by which the food is taken hastily into
the paunch and afterwards sent back to the mouth in detached portions
for farther mastication, is called _rumination_ or _chewing
the cud_, and those species which perform it are thence called
_ruminating_ animals. Sheep and cows may be seen lying ruminating
in pasture-fields after having cropped as much herbage as fills the
paunch; and feeding is thus rendered to them a source of prolonged
enjoyment.
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