Burrowing-snakes live chiefly underground, and often have the visual
organ atrophied in consequence, as in _Typhlops_; all the Typhlopidæ,
Glauconiidæ, and Uropeltidæ, belong to this category; the Viperid
_Atractaspis_ is also a burrowing type.
Tree-snakes spend the greater part of their life on bushes or trees.
_Corallus_ among the Boidæ, _Dendrophis_ and _Dendraspis_ among the
Colubridæ, _Atheris_ and various species of _Lachesis_ among the
Viperidæ, may be quoted as examples.
Of Water-snakes, some are exclusively aquatic, like the marine
Hydrophiinæ and the typical Acrochordinæ (_Acrochordus_, _Chersydrus_)
and Homalopsinæ (_Hipistes_, _Herpeton_). _Chersydrus_ and _Hipistes_
occur in the sea as well as in fresh water. Many species of
_Tropidonotus_ (_T. tessellatus_ and _T. viperinus_ in Europe), as well
as the genera _Helicops_, _Grayia_, _Boulengerina_, etc., among the
Colubridæ, _Eunectes_ among the Boidæ, _Ancistrodon piscivorus_ among
the Viperidæ, are chiefly but not exclusively aquatic.
Our _Tropidonotus natrix_ stands between the Ground-snakes and the
Water-snakes; Boas and Pythons are as much Water-snakes as Tree-snakes.
As shown by these and many other examples which might be given, a
division into categories cannot always be applied with precision, nor
does it convey an expression of the natural relationships of the
species, as was believed by many systematists of the last century, who
appealed to such adaptations for the definition of families.
A vertical pupil denotes more or less nocturnal habits. Nevertheless our
European Vipers, which are provided with such a contractile pupil, are
far from exclusively nocturnal, delighting to bask in the sun, and
pairing and feeding in the day-time. The Boidæ appear to be more
nocturnal, but no snake is known to be absolutely so, and the two
species of _Coluber_ which have been found living in perfect darkness in
limestone caves in the Malay Peninsula and China, where they feed
chiefly on bats, occur also outside the caves, and probably never breed
in them.
It is often stated in books that the organs of locomotion for the
exceedingly elongate body of snakes are the ribs, and these creatures
have even been compared to Centipedes. This statement is no doubt true
to a certain extent for slow locomotion on uneven ground, when the ribs
and the corresponding ventral shields afford a point of support; but it
does not account for the rapid movements, as when a snake darts like an
arrow in pursuit of its prey or to escape from an enemy. Besides, the
winding motions are not different from those of a Slow-worm or
Glass-snake, in which, encased as they are in a bony armour, the ribs
cannot come into play at all. The action of the muscles alone is quite
sufficient to account for the reptation of snakes, without the ribs
having to play an essential part.
Public-domain text, read in full here on John Shaqi.
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