A history of land mammals in the western hemisphereScott, William Berryman
History
A history of land mammals in the western hemisphere
Scott, William Berryman
Mammals; Mammals, Fossil
by great thickening of the cranial bones, as is true of elephants and
rhinoceroses, and in them sufficient surface for attachment is afforded
to the muscles without the development of a crest.
[Illustration: FIG. 8.—Skull of Wolf, top view. _P.Mx._, premaxillary.
_Na._, nasal. _Ma._, malar or jugal. _L._, lachrymal. _Fr._, frontal.
_Sq._, squamosal. _Pa._, parietal. _S.O._, supraoccipital.]
[Illustration: FIG. 9.—Skull of Wolf, view of base. _P.Mx._,
premaxillary. _Mx._, palatine process of maxillary. _Pl._, palatine.
_Fr._, frontal. _Pt._, parietal. _Ma._, malar or jugal. _Sq._, glenoid
cavity of squamosal. _B.S._, basisphenoid. _B.O._, basioccipital. _Ty._,
tympanic (auditory bulla). _p.oc._, paroccipital process. _con._,
occipital condyle. _S.O._, supraoccipital.]
The structure of these cranial bones, more particularly of the parietals,
is subject to important changes; in most mammals they are of moderate
thickness and have dense layers, or “tables,” forming the outer and
inner surfaces and, between these, a layer of spongy bone. In many large
mammals, however, especially those which have heavy horns or tusks, the
cranial bones become enormously thick and the spongy layer is converted
into a series of communicating chambers, or _sinuses_, the partitions
between which serve as braces, thus making the bone very strong in
proportion to its weight. Sinuses are very generally present in the
frontals and communicate by small openings with the nasal passage, even
in genera of moderate size and without horns or tusks. The frontals
form the roof of the eye-sockets, or _orbits_, and usually there is
a projection from each frontal, which marks the hinder border of the
orbit and is therefore called the _postorbital_ process. The roof of the
facial region is made by the _nasals_, which are commonly long and narrow
bones, but vary greatly in form and proportions in different mammals;
in those which have a proboscis, like tapirs and elephants, or a much
inflated snout, such as the Moose (_Alce_) or the Saiga Antelope (_Saiga
tatarica_) the nasals are always very much shortened and otherwise
modified in form.
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