What is protoplasm? In its substance it is a nitrogenous carbon compound,
differing only from other similar compounds of the albuminous family of
colloid by the extremely complex composition of its atoms. It consists
of five elements, and its average composition is said by chemists to be
52·55 per cent. carbon, 21·23 oxygen, 15·17 nitrogen, 6·7 hydrogen, 1·2
sulphur. Its peculiar qualities, therefore, including life, are not the
result of any new and peculiar atom added to the known chemical compounds
of the same family, but of the manner of grouping and motions of these
well-known material elements. It has in a remarkable degree the faculty
of absorbing water, so that its molecules seem to float in it in a
condition of semi-fluid aggregation, which seems to be necessary for the
complex molecular movements which are the cause or accompaniment of life.
Thus, many seeds and animalculæ, if perfectly dry, may remain apparently
as dead and as unchanging as crystals, for years, or even, as in the case
of the mummy wheat, for centuries, to revive into life when moistened.
But in addition to those material qualities in which protoplasm seems
to differ only from a whole group of similar compounds of the type of
glycerine, by the greater complexity and mobility of its molecules, it
has developed the new and peculiar element which is called life. Life
in its essence is manifested by the faculties of nutrition, sensation,
movement, and reproduction.
As regards nutrition there is this essential difference between living
and non-living matter. The latter, if it feeds and grows at all, does
so only by taking on fresh molecules of its own substance on its outer
surface, as in the case of a small nucleus-crystal of ice in freezing
water. If it feeds on foreign matter and throughout its mass, it does
so only in the way of chemical combination, forming a new product.
Living matter, on the other hand, feeds internally, and works up
foreign substances, by the process we call digestion, into molecules
like its own, which it assimilates, rejecting as waste any surplus or
foreign matter which it cannot incorporate. It thus grows and decays
as assimilation or waste preponderates, remaining always itself. The
distinction will be clear if we consider what happens when water rusts
iron. In a certain sense the iron may be said to eat the oxygen, reject
the hydrogen, and grow, or increase in weight by what it feeds on; but
the result is not a bigger piece of iron, but a new substance, rust,
or oxide of iron. That living matter should feed internally is not
so wonderful, for its semi-fluid condition may well enable foreign
molecules to penetrate its mass and come in contact with its own interior
molecules; but it is an experience different from anything known in
the inorganic world that it should be able to manufacture molecules
of protoplasm like its own out of these foreign molecules, and thus
grow by assimilation. For instance, when amœbæ, bacteria, and other
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