A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
_Number and Arrangement of the Compartments._--I have already stated
that the shell, in every one of the Balanidæ, consists of eight, six,
or four compartments, or of all fused together into a single piece;
and that the compartments themselves are all constructed on the three
simple patterns of which woodcuts (figs. 1, 2, 3) have been given.
They are arranged in a certain definite order. The type arrangement is
found amongst the Chthamalinæ, as might have been expected, inasmuch
as this sub-family is so closely related to the ancient genera
Pollicipes and Scalpellum, whence all the Thoracic Cirripedia may be
said to radiate. In Octomeris (fig. 4) the type-arrangement of the
compartments, eight in number, is well shown; the rostrum and carina
resemble each other, and have alæ on both sides, and therefore are
overlapped on both sides: the rostro-lateral compartments have radii
on both sides, and therefore overlap the adjoining compartments on both
sides; the lateral and carino-lateral compartments have radii on their
carinal, and alæ on their rostral sides; and therefore overlap on one
side, and are overlapped on the other side. Now the shell of every
other sessile Cirripede differs, I believe, from that of Octomeris,
only in the fusion together or abortion of some of the eight normal
compartments: in one genus, however, Catophragmus, outer whorls of
small compartments, arranged like the lower valves in the capitulum
of Pollicipes, are superadded. The genus Chthamalus (fig. 5) differs
from Octomeris only in the carino-lateral compartments being aborted,
(as will presently be discussed), and hence has six compartments.
Chamæsipho (fig. 6) differs from Chthamalus only in the rostro-lateral
and lateral compartments being fused together; and hence has only four
compartments. In Balanus (fig. 7) and the whole sub-family of the
Balaninæ, the rostrum is compounded of the true rostrum, as seen in the
type Octomeris, and of the two rostro-lateral compartments; hence this
compounded rostrum has radii instead of alæ on both sides, and there
are only six compartments. Tetraclita (fig. 8) and Elminius differ
from Balanus only in having the carino-lateral compartments absent, and
probably aborted; hence there are only four compartments. Lastly, in
Pyrgoma, all the compartments are blended together into a single piece.
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