A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
_Anomalies and exceptions._--In Tetraclita (Pl. 10, fig. 1 _g_, 1 _h_)
from the branching of the longitudinal septa, several irregular rows
of tubes are formed. In certain varieties of _Balanus balanoides_
(Pl. 7, fig. 2 _b_), and in _B. cariosus_ (Pl. 7, fig. 3 _b_), slight
branching ridges on the internal surface of the walls, seem to
answer to the longitudinal septa, and produce, during the downward
growth of the shell, extremely irregular cells, and short tubes. In
_Balanus vinaceus_ (Pl. 2, fig. 7 _d_), the internal lamina, instead
of being solid, as in every other species, is left cancellated, and
thus betrays, much more plainly than usual, its origin in the united
denticuli of the adjoining longitudinal septa. In _Balanus porcatus_,
between the main longitudinal septa, there are (Pl. 6, fig. 4 _e_)
what may be called rudimentary and disconnected longitudinal septa. In
Coronula and its allies (Pl. 16, fig. 6, and Pl. 17, fig. 4 _c_) it
is the outer lamina of the compartment which is anomalous; for in the
two or three lower zones of growth, it forms only a ledge on each side
of the longitudinal septa; which ledges, higher up, become confluent,
and so form an ordinary outer lamina. In Coronula, also, the wall of
each compartment (see transverse section, Pl. 16, figs. 5, 7) is very
remarkable from being deeply folded, the folds being on their internal
faces firmly calcified together, and on their external faces closely
pressed together (often with a neatly serrated suture), so that the
whole nature of the shell might be, as has happened, easily quite
misunderstood; and the walls be considered as very thick, instead of
being, as is really the case, very thin. In Chelonobia (Pl. 15, fig.
1), however, the walls are truly of such great thickness, that the
nature of the relative parts might likewise be misunderstood; in this
genus the ovarian tubes enter the walls, extending up between the
longitudinal septa, or, as they may here be more naturally called,
the radiating septa. I will specify a few more peculiarities worthy
of remark:--in some species of the sub-genus Acasta, clefts are left,
covered only by membrane, on the lines of suture (Pl. 9, figs. 7 _a_,
8 _a_), between the compartments, just above the basis; and in other
species the basis is perforated by numerous membrane-covered, minute
orifices. In Platylepas, each compartment has one deep inward fold (Pl.
17, fig. 1), somewhat analogous to the three folds in Coronula; this
fold is produced into an internal midrib, supporting and rendering
convex the membranous basis; in this genus, also, the rostrum, owing
to its midrib, is generally thrust a little on one side, and the shell
thus rendered asymmetrical. In _Chamæsipho scutelliformis_ the shell
is symmetrically perforated (Pl. 19, fig. 4 _a_) by four apertures.
Lastly, in _Chthamalus Hembeli_ and _intertextus_, after a certain age,
the basal edges of the walls become inflected, and continue to grow
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