An introduction to the study of fishesGünther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Science
An introduction to the study of fishes
Günther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Fishes; Ichthyology
The chain of flat bones which, after the removal of the temporal
muscles, appear arranged within the inner concavity of the præoperculum
(Fig. 24), are comprised with the latter under the common name of
_mandibulary suspensorium_. They connect the mandible with the
cranium. The uppermost, the _epitympanic_ or _hyomandibular_ (23), is
articulated by a double articulary head with the mastoid and posterior
frontal. Another articulary head is destined for the opercular joint.
The _mesotympanic_ or _symplectic_ (31) appears as a styliform
prolongation of the lower part of the hyomandibular; is entirely
cartilaginous in the young, but nearly entirely ossified in the adult.
The position of this bone is noteworthy, because, directly inwards of
its cartilaginous junction with the hyomandibular, there is situated
the uppermost piece of the hyoid arch, the stylohyal. The next bone of
the series is the _pretympanic_ or _metapterygoid_ (27), a flat bone
forming a bridge towards the pterygoid, and not rarely absent in the
teleosteous sub-class. Finally, the large triangular _hypo-tympanic_ or
_quadrate_ (26) has a large condyle for the mandibulary joint.
The palatine arch (Fig. 26) connects the suspensorium with the
anterior extremity of the skull, and is formed by three bones: the
_entopterygoid_ (25), an oblong and thin bone attached to the inner
border of the palatine and pterygoid, and increasing the surface of
the bony roof of the mouth towards the median line; it constitutes
also the floor of the orbit. The _pterygoid_ (24) (or _os transversum_)
starts from the quadrate, and is joined by suture to the _palatine_,
which is toothed, and reaches to the vomer and anterior frontal.
In the occipital region there are distinguished the _basi-occipital_
(5), readily recognised by the conical excavation corresponding and
similar to that of the atlas, with which it is articulated through
the intervention of a capsule filled with a gelatinous substance (the
remains of the notochord); the _exoccipitals_ (10), articulated, one on
each side, to the basi-occipital, and expanding on the upper surface of
that bone, so as to meet and support the spinal column; a superficial
thin lamella (13), suturally connected with the exoccipitals,
not constant in fishes, and erroneously believed by Cuvier to be
the _petrosal_ (_os petrosum_) of higher animals; further, the
_paroccipitals_ (9), which are wedged in between the exoccipitals and
_supraoccipital_. This last bone (8) forms the key of the arch over the
occipital foramen, and raises a strong high crest from the whole length
of its mesial line; a transverse supraoccipital ridge, coming from each
side of the base of this spine runs outwards laterally to the external
angles of the bone. The supraoccipital separates the parietals, and
forms a suture with the frontals.
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