Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
Referring again to fig. 37 as a starting-point, imagine the pith in the
centre to spread in a star-shaped form till the points of the star
touched the edges of the ring, and thus to break the wood ring into
groups. A stage in this process (which is not yet completed) is shown in
fig. 41, while in fig. 42 the wood and bast groups are entirely distinct.
In the flowering plants the cells of the endodermis are frequently poorly
characterized, and the pith cells resemble those of the cortical ground
tissue, so that the separate groups of wood and bast (usually known as
“vascular bundles”, in distinction from the “steles” of fig. 40) appear
to lie independently in the ground tissue. These strands, however, must
not be confused with steles, they are only fragments of the single
apparently broken up stele which runs in the stem.
[Illustration: Fig. 42.—Monostele in which the Pith has invaded all the
Tissues as far as the Endodermis, and broken the Wood and Phloem up
into Separate Bundles. These are usually called “vascular bundles” in
the flowering plants]
[Illustration: Fig. 43.—Showing actively growing Zone _c_ (Cambium) in
the Vascular Bundles, and joining across the ground tissue between them]
The vascular bundle, of all except the Monocotyledons, has a potentiality
for continued growth and expansion which places it far above the stele in
value for a plant of long life and considerable growth. The cells lying
between the wood and the bast, the soft parenchyma cells always
accompanying such tissues, retain their vitality and continue to divide
with great regularity, and to give rise to a continuous succession of new
cells of wood on the one side and bast on the other; see fig. 33, _c_,
_b_. In this way the primary, distinct vascular bundles are joined by a
ring of wood, see fig. 43, to which are added further rings every season,
till the mass of wood becomes a strong solid shaft. This ever-recurring
activity of the cambium gives rise to what are known as “annual rings” in
stems, see fig. 44, in which the wood shows both primary distinct groups
in the centre, and the rings of growth of later years.
Cambium with this power of long-continued activity is found in nearly all
the higher plants of to-day (except the Monocotyledons), but in the fern
and lycopod groups it is in abeyance. Certain cases from nearly every
family of the Pteridophytes are known, where some slight development of
cambium with its secondary thickening takes place, but in the groups
below the Gymnosperms cambium has almost no part to play. On the other
hand, so far back as the Carboniferous period, the masses of wood in the
Pteridophyte trees were formed by cambium in just the same way as they
are now in the higher forms. Its presence was almost universal at that
time in the lower groups where to-day there are hardly any traces of it
to be found.
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