Appletons' Popular Science Monthly, December 1898: Volume LIV, No. 2, December 1898Various
Science
Appletons' Popular Science Monthly, December 1898: Volume LIV, No. 2, December 1898
Various
Science -- Periodicals; Technology -- Periodicals
In general it is to be said that stamens and pistils may reach
functional maturity in darkness or diffuse light, and if pollination
is provided for, seed and fruit formation may ensue.
The diminution of light has the effect of transforming inflorescences
into leafy shoots in some instances, however. The more common
reaction consists of alterations in the size, form, and color
of the perianth, and greater changes are induced in the petals
than in the sepals. The corolla shows greater decrease in size in
_Melandryum_ and _Silene_, in diffuse light, though the relative
form is maintained. The writer has obtained most striking results
from growing flowers of _Salvia_ (sage) in a dark chamber, inclosing
the inflorescence only. In the normal flower the irregular scarlet
corolla attains three times the length of the calyx, and two stamens
extrude from under the upper lip. When grown in darkness, the corolla
with the adherent stamens measure about three millimetres in length,
or one twelfth the normal, and are scarcely more than half the size
of the calyx, which is but two thirds the size of similar organs
grown in the light. The color is entirely lacking from the corolla,
and is found only along the veins of the calyx.
In other instances in which the corolla is composed of separate
members, an unequal reaction is exhibited. The corolla of nasturtium
(_Tropæolum majus_) consists of five approximately equal petals.
Flowers of this species grown in darkness show one of nearly normal
stature, two of reduced size, while the remaining two take the form
of club-shaped bracts.
The diminished size of the perianth of cleistogamous flowers of such
types as the violet is due directly to the action of diminished light
upon the hidden or inclosed flower.
The influence of light upon the structure, reproductive processes,
and distribution of the lower forms brings about the most widely
divergent reactions, which can not be described here.
The distribution and color of marine algæ depend upon the depth of
the water and the consequent intensity of the light. This gives
rise to distinct zones of aquatic vegetation. Thus in one series of
surveys the _littoral_ zone, the beach area covered at high water
and exposed at low water, was found to furnish proper conditions
for green, brown, and red algæ. The _sublittoral_ zone, extending
to a depth of forty metres, is furnished with red algæ, increasing
in number with the depth, and the brown algæ disappear; while
the _elittoral_ zone, from forty to one hundred and ten metres,
is inhabited by red algæ alone. The number of species of vegetal
organisms below this depth is extremely small. An alga (_Halosphæria
viridis_) has been brought up from depths of one thousand to two
thousand metres.
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