Aspects of plant life; with special reference to the British floraPraeger, R. Lloyd (Robert Lloyd)
Science
Aspects of plant life; with special reference to the British flora
Praeger, R. Lloyd (Robert Lloyd)
Plants -- Great Britain
But it is in the flower itself that we find the most ingenious
arrangements to encourage useful and discourage useless visitors, to
assist the former to pollinate the flower, and while offering nectar to
the welcome guest to deny it to the unwelcome. The first stage in this
specialization is that the flower, instead of having its axis vertical,
and facing the sky, is turned on its side by the curving of its stalk,
and looks out horizontally. The effect of this is to cause a flying
insect on approaching the flower to alight in a particular
position--namely, on the lowest petal. Following on the adoption of this
attitude the next stage in development is seen in the parts of the
flower beginning to alter their shape and position relative to each
other and often also their colour. Thus, beginning with a quite regular
flower, we can arrange a series showing more and more asymmetry. The
tendency is generally for the lowest petal to become enlarged and often
conspicuously marked, providing a broad, convenient platform on which
insects may alight, while the remainder form walls and roof, protecting
the important parts within and by their shape, which is often narrowed
and tubular behind, barring access to all but chosen visitors. To find a
full series illustrating these transformations we do not need to go to
plants widely separated in their affinities. In the Buttercup order
(_Ranunculaceæ_) alone every gradation may be found. The flowers of the
Buttercups themselves are upright and quite regular. In the Larkspur
(_Delphinium_) the flower is turned on its side, and a puzzling
combination of coloured sepals and petals--five bright blue unequal
sepals and a single large purplish petal of peculiar shape with a long
hollow spur behind--produces a quite irregular blossom. The process is
carried farther again in the Monkshood (_Aconitum_), in whose well-known
blue flower the sepals and petals combine to produce a strikingly
irregular blossom, with the upper sepal arching over into a great hood
protecting the rest of the flower. In such irregular flowers the
essential parts--the pollen-producing and pollen-receiving portions, or
stamens and stigma--also alter their position and form, and are so
placed that an insect, visiting the flower to obtain nectar (which is
generally stored at the back, well out of the way), must of necessity
receive pollen on its body, and probably deposit pollen on the stigma.
To describe the variety and ingenuity of these devices as found in
different flowers might well occupy several chapters, and only one or
two examples can be quoted here; familiar wild flowers are chosen, and
the reader should examine them for himself to understand their
structure. In the well-known Pea type, one great petal arches over the
flower; two narrow ones stand one on either side; the remaining two
stand on edge below, with their margins in contact, enclosing the
stamens and pistil. An insect visiting the flower alights naturally on
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