Aspects of plant life; with special reference to the British floraPraeger, R. Lloyd (Robert Lloyd)
Science
Aspects of plant life; with special reference to the British flora
Praeger, R. Lloyd (Robert Lloyd)
Plants -- Great Britain
Only one more instance can be referred to, which can be tested by the
reader any summer day wherever any of our native Orchids grow. In
these, the most highly specialized of all plant groups as regards
pollination by insects, the general arrangement of the flower is often
somewhat similar in a general sense to the last case; but here the
sepals and petals which between them form the platform, tube, sides, and
roof of the flower, are all separate and often differently and
elaborately coloured. The essential organs are greatly modified and
hardly recognizable at first. There is only one stamen, producing two
clusters of pollen, which are embedded in the roof of the flower. Each
possesses a slender stalk which terminates in a little sticky disc which
projects from the general surface. The pollen grains are held together
in a mass by fine threads, and the whole with its stalk--the
_pollinium_--resembles a lemonade bottle in shape. The stigma is also
embedded, forming a sticky surface in the roof of the flower behind the
stamen. When an insect inserts its head into the flower, its forehead
comes in contact with the sticky ends of the pollinia, which adhere, so
that on leaving the flower the insect flies away with the pollen
sticking to its forehead like two little horns. And now a remarkable
thing happens. The stalks of the pollinia, drying rapidly in the air,
contract unequally, and become curved, so that the pollinia bend forward
into a horizontal position. When the insect visits another flower and
thrusts in its head, the pollen consequently comes in contact with the
sticky stigmatic surface farther down the tube, and cross-pollination is
effected.
In the cases of many of these highly specialized flowers, one is no less
struck with the perfection of the arrangements made for preventing
self-pollination, than those adapted to securing cross-pollination. But
in a few, on the contrary, self-pollination is specially arranged for.
It must be pointed out that the insects which pollinate these
specialized flowers have in many cases acquired modifications in their
structure corresponding to the modifications in the flowers which they
frequent. In the more specialized forms, indeed, plant and animal have
become entirely dependent on each other; the plants would become extinct
in the absence of the special insects through whose agency they are able
by pollination to produce fertile seed; and the insects would likewise
die out if the flowers to whose nectar and pollen they look for food
were not available.
Public-domain text, read in full here on John Shaqi.
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