Indeed, there seem to be only two tribes which have any use for
wing-claws during nestling life. One of these is represented by the
gallinules, that is to say, the coots, and water-hens, and their kind.
You may test this whenever you have the good fortune to capture a young
water-hen. Place him outside the nest, and especially if it happens to
be a little raised, you will see him make his way back, using feet,
wing-claws, and beak. His wings, it will be noticed, at this stage are
used as fore-legs. The other tribe is represented by that strange bird
the hoatzin of the Amazon. Here the two claws are really large, and
they play a quite important part in his early life.
For the young hoatzin is hatched in a nursery--a crude nest of
sticks--placed on the boughs of a tree overhanging the water. As soon
as hatched he begins to climb about the branches. Should he fall, by
some mischance, into the water, he promptly swims to the bank; and by
the aid of his long first finger, and wing-claws, and his huge feet,
soon climbs back. But the most wonderful part of his story is yet to
come.
[Illustration: _Grouse_]
So long as these youngsters can only scramble about they are in
constant jeopardy. A wing-surface at least big enough to break the
force of a fall is an urgent necessity. And so the growth of the
quill-feathers is, so to speak, pushed forward with all possible speed.
But if all the feathers grew at the same rate, there would speedily
come a time when the outermost feathers would make the claw at the
end of the finger useless, while the wing-surface, as a whole, would
be insufficient. To obviate this difficulty, the development of the
outermost feathers is held in abeyance till the inner feathers of
the hand, and the outermost of the fore-arm, have grown big enough
to suffice to break the force of the fall. As soon as this stage
is arrived at, the outermost quills, whose growth has been held in
abeyance, rapidly develop; the finger decreases in length, and its claw
disappears, while that of the thumb soon follows suit. And thus it
comes about that the hand, in the nestling, is relatively much longer
than in the adult. But in its mid-period it may be taken to represent
the adult stage of the wing of the ancient Archæopteryx. This bird
could have been but a poor flier, and probably during the time it was
moulting its quills it was absolutely flightless, so that it needed a
permanent finger-tip, and claw, beyond the margin of its wing-surface.
Public-domain text, read in full here on John Shaqi.
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