This matter of "moulting," by the way, needs, at least, passing
comment. All birds renew their plumage at least once: the body plumage
often twice in the year. The old feathers fall out, and their places
are taken by new ones. But their growth is slow. In geese and ducks,
and some other birds, the wing-quills are moulted all at once, so that
flight, for a week or two, is impossible. But they can escape from
their enemies while thus at a disadvantage, by taking to the water. In
all other birds the quills are moulted, and renewed, in pairs: so that
at no time are they left flightless.
But this by the way. Let us revert, for a moment, to the hoatzin's
wing. The appearance of the outermost quills of the hand, it will be
remembered, is delayed till the inner feathers have grown long enough
to "flutter," at least, for a short distance, then the growth of the
complete series proceeds apace. This has been called an "Adaptation"
to enable these youngsters, active from the moment they leave the egg,
to move about in comparative safety. But it is more than this. It is a
survival of an ancient order of things which takes us back to the first
known birds.
This is certainly a very remarkable feature, but it gains an added
interest from the fact that it has a parallel in the history of the
development of the wing in the game-birds. If you look carefully at
the downy chicks of the pheasant, or even at barn-door fowls, you
will remark that the wing-quills develop with surprising rapidity: so
that they have feathered wings while the rest of the body is still
down-covered. This enables them the more easily to escape prowling
foxes and other enemies. In young ducks exactly the opposite condition
obtains, the body is fully feathered long before the feathers of the
wings appear. And this because they do not need to fly when danger
threatens, but take to the water instead. But to return to the chicks
of the pheasant. The wing of the chick develops at a very rapid rate.
Within a few hours after hatching, the first traces of the coming
flight feathers can be seen, and presently a large wing is covering
each side of the tiny body. At this stage many often die. The wings,
which can then be examined at leisure, reveal an extremely interesting
condition. For they repeat the features which obtain in the wing of the
nestling hoatzin: inasmuch as the outermost quills are also, as yet,
non-existent; and there is a free finger-tip. But it is not nearly so
long as in the hoatzin, and there is no terminal claw. Surely, from
this, we may infer that the delayed development of the outer quills is
a survival of a time when the ancestors of the pheasant were arboreal,
and hatched their young in trees. Otherwise all the wing-quills should
develop at the same time, and at the same rate. Here, then, is another
instance of what can be learned of the past history of a bird by a
careful scrutiny of the nestling. Sometimes we shall find our evidence
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