If we could stretch an apparently impervious membrane, like the inner
white skin just inside an eggshell, or a piece of parchment, and so form
a wall through the middle of a glass box, and then pour into one of the
compartments pure water and in the other a mixture of water and
molasses, a very curious result would follow within a comparatively
short period. We should find that presently there would be a gentle
filtering of the water through the membrane toward the molasses water,
and similar gentle current in the other direction. In other words,
fluids of different density, if separated by a membrane, tend to
equalize each other. This equalization may not be very rapid, and at
first it will be more speedy from the less dense to the more dense, but
eventually it will make the different fluids of a common density. This
purely mechanical property of the equalization of fluids separated by a
membrane is known as _osmosis_, and it is upon the possession of the
equipment necessary for this that roots depend for getting food and
water from the soil.
In our discussion of roots in Chapter I, we found that they end in very
fine subdivisions, which are themselves split up into practically
invisible root hairs. These root hairs are the only way that roots can
absorb the food and water in the soil, and they are able to do this
because they are provided with a membrane which permits osmosis to act
between the solution inside the root hair and the water in the soil. The
solution in the root hair is mostly a sugary liquid, some of that
surplus sugar made in the leaves, and it is denser than the soil water,
so there is apparently nothing to prevent an equalization of the liquids
on different sides of the membrane. If this actually happened, as it
would in the case of the simple experiment noted above, then roots would
exchange a fairly rich sugary liquid for a much more watery one, and we
should find that plants did not get their food from the soil, but really
have it drained away from them by osmosis. But nature has a cunning
device for stopping such robbery, which is prevented by the membranes of
root hairs being only permeable to the extent of letting water _in_, not
permeable enough to allow sugar to escape. As we have seen, osmosis is
a purely mechanical process which, if left to operate without
interference, would not aid but injure the plant. Surely, nothing with
which plants are provided is so important to them as this delicate
membrane of the root hairs which, while allowing osmosis to act in a
one-sided fashion, preserves to the plant the sugary liquid that alone
makes the absorption of soil water possible.
Public-domain text, read in full here on John Shaqi.
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