If you will turn back for a moment to the beginning of the description
of how plants get their food, you will find that in osmosis the weaker
liquid tends to permeate the denser one more rapidly than the denser one
does the weaker. As we have just seen, the sugary liquid in the root
hairs is denser than the soil water outside, and, furthermore, _none of
it is allowed to escape_. This comparatively greedy process of taking
everything and giving nothing results in a constant flow of soil water
into the root hairs. When the flow of liquids in osmosis is not at once
equalized, a gentle pressure is brought to bear to make them so. This is
what is called _osmotic pressure_, and it is this pressure that forces
the absorbed liquid through the roots and part way up the trunk of even
the tallest trees. While we have just said it is a gentle pressure, that
is true only in the case where the osmosis has free play, and the
pressure is stopped with the perfect mixing of the two liquids. But what
if they can never mix? What may not the accumulated osmotic pressure
amount to in such a one-sided process as goes on in root hairs with
everything coming in and nothing going out. Cut-off stems, with a
pressure gauge attached to them, indicate that in some plants the
pressure is from 60 up to 170 pounds!
Another result of this pressure is that it keeps leaves and the fleshy
stems of plants in their ordinary position. The actual solid part of
nearly all leaves is scarcely 5 per cent of their bulk and all the rest
is water. The constant pressure of this water from the roots is
sufficient to keep leaves comparatively stiff and rigid, how stiff is
quickly realized if the pressure stops and the leaf wilts or withers.
Sometimes this osmotic pressure, particularly during rainy weather,
becomes so great as to cause injury to the plant, the splitting of
tomatoes and occasionally of plums, being due to it. This osmotic
pressure, together with the extra pull given by the leaves, is
sufficient to account for the rise of water to the tops of the tallest
trees. The tallest trees in the world are certain kinds of blue gum in
Australia which frequently reach a height exceeding 300 feet. What the
combined osmotic pressure and leaf pull must be to carry such a heavy
thing as water to such a great height is easier to imagine than to
calculate.
Public-domain text, read in full here on John Shaqi.
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