Compare the hinder grasping (so-called) hand
of the Gorilla with this.
At first sight there is a great difference, for the great toe and its
metatarsal bone form _an angle_ with the bones of the other toes and
their metatarsals. Instead of the toes and their slender bones being
parallel and fixed in this position, the great toe of the Gorilla has
a power of moving so as to cross the foot more or less below, as the
human thumb can cross the palm. It has also the capacity of being
stretched out from the foot, so that its movements greatly resemble
those of a thumb. In fact, we want a word to express a toe-thumb.
On examining the foot carefully, it will be found that each of its
bones may be compared and identified in position and office with the
same in man. There is a heel-bone with a great projection behind, for
the fixing on of the Achilles tendon, and this is jointed on to a bone
above, like the human die-bone or astragalus, and to one in front,
like the cuboid. The astragalus resembles that of man, but the upper
and outer surfaces on which the lower ends of the leg-bones move, are
slightly different, so as to admit of greater turning in of the ankle.
The wedge-shaped bones are there, and the inner one, with its joint for
the slender bone of the great toe, is shorter and broader than in man,
so as to allow of great movement of the toe-thumb in front of it. The
slender bones, or metatarsals, are larger and longer, but their shape
and direction, with the exception of the first, are singularly like
those of man. As a whole the foot of the Gorilla, for thus it must be
called, is broader in front of the ankle-bones and longer everywhere
than in us, but it has a sideway and almost club-foot look about it;
its position is “turned in,” like the foot of a young child before it
walks. This is owing to the conformation and easy jointing of the bones
of the ankle and foot, and also to the action of a front muscle of the
leg which pulls the very movable bones inwards. The structures allow
of a very ready turning in of the ankle and foot, and such as would
render climbing easy with the aid of the toe-thumb; but they evidently
interfere with the steadiness in walking. It is a huge foot, and it
is only half an inch or so shorter than the leg below the knee; it is
unwieldy as a foot, but is a capital foot-hand, which cannot readily
have its toes stretched out straight, for their usual position is that
of being slightly bent in the direction of the sole.
Public-domain text, read in full here on John Shaqi.
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