Let us assume, to begin with, the Darwinian theory of insensible variations, and suppose the occurrence of small differences due to chance, and continually accumulating. It must not be forgotten that all the parts of an organism are necessarily coördinated. Whether the function be the effect of the organ or its cause, it matters little; one point is certain--the organ will be of no use and will not give selection a hold unless it functions. However the minute structure of the retina may develop, and however complicated it may become, such progress, instead of favoring vision, will probably hinder it if the visual centres do not develop at the same time, as well as several parts of the visual organ itself. If the variations are accidental, how can they ever agree to arise in every part of the organ at the same time, in such way that the organ will continue to perform its function? Darwin quite understood this; it is one of the reasons why he regarded variation as insensible.[29] For a difference which arises accidentally at one point of the visual apparatus, if it be very slight, will not hinder the functioning of the organ; and hence this first accidental variation can, in a sense, _wait for_ complementary variations to accumulate and raise vision to a higher degree of perfection. Granted; but while the insensible variation does not hinder the functioning of the eye, neither does it help it, so long as the variations that are complementary do not occur. How, in that case, can the variation be retained by natural selection? Unwittingly one will reason as if the slight variation were a toothing stone set up by the organism and reserved for a later construction. This hypothesis, so little conformable to the Darwinian principle, is difficult enough to avoid even in the case of an organ which has been developed along one single main line of evolution, _e.g._ the vertebrate eye. But it is absolutely forced upon us when we observe the likeness of structure of the vertebrate eye and that of the molluscs. How could the same small variations, incalculable in number, have ever occurred in the same order on two independent lines of evolution, if they were purely accidental? And how could they have been preserved by selection and accumulated in both cases, the same in the same order, when each of them, taken separately, was of no use?