Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
We have already seen that before the subject was taken up by Weismann
the difference between acquired and congenital characters in respect to
transmissibility was generally taken to be one of degree; not one of
kind. It was usually supposed that acquired characters, although not so
fully and not so certainly inherited as congenital characters,
nevertheless were inherited in some lesser degree; so that if the same
acquired character continued to be successively acquired in a number of
sequent generations, what was at first only a slight tendency to be
inherited would become by summation a more and more pronounced tendency,
till eventually the acquired character might become as strongly
inherited as a congenital one. Or, more precisely, it was supposed that
an acquired character, in virtue of such a summation of hereditary
influence, would in time become congenital. Now, if this supposition be
true, it is evident that more or less assistance must be lent to
natural selection in its work of evolving adaptive modifications[28].
And inasmuch as we know to what a wonderful extent adaptive
modifications are secured during individual life-times--by the direct
action of the environment on the one hand, and by increased or
diminished use of special organs and mental faculties on the other--it
becomes obvious of what importance even a small measure of
transmissibility on their part would be in furnishing to natural
selection ready-made variations in required directions, as distinguished
from promiscuous variations in all directions. Contrariwise, if
functionally-produced adaptations and adaptations produced by the direct
action of the environment are never transmitted in any degree, not only
would there be an incalculable waste, so to speak, of adaptive
modifications--these being all laboriously and often most delicately
built up during life-times of individuals only to be thrown down again
as regards the interest of species--but so large an additional burden
would be thrown upon the shoulders of natural selection that it becomes
difficult to conceive how even this gigantic principle could sustain it,
as I shall endeavour to show more fully in future chapters. On the other
hand, however, Weismann and his followers not only feel no difficulty in
throwing overboard all this ready-made machinery for turning out
adaptive modifications when and as required; but they even represent
that by so doing they are following the logical maxim, _Entia non sunt
multiplicanda praeter necessitatem_--which means, in its relation to
causality, that we must not needlessly multiply hypothetical principles
to explain given results. But when appeal is here made to this logical
principle--the so-called Law of Parsimony--two things are forgotten.
Public-domain text, read in full here on John Shaqi.
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