Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
this is impossible in the case of inherited characters which are also
_adaptive_ characters. How then does it fare with the case of inherited
characters which are not also adaptive? Merely that this case is met by
another and sequent assumption, which constitutes an integral part of
the Neo-Darwinian creed--namely, that in nature there _can be no such
characters_. Seeing that natural selection is taken to be the only
possible cause of change in species, it follows that all changes
occurring in species must necessarily be adaptive, whether or not we are
able to perceive the adaptations. In this way apparently useless
characters, as well as obviously useful ones, are ruled out of the
question: that is to say, _all_ hereditary characters of species in a
state of nature are _assumed_ to be due to natural selection, and then
it is demanded that the validity of this assumption should be disproved
by anybody who doubts it. Yet Weismann himself would be unable to
suggest any conceivable method by which it can be disproved among
species in a state of nature--and this even supposing that the
assumption is entirely false[30].
[30] In subsequent chapters, especially devoted to the question
(i.e. Section II), the validity of this assumption will be
considered on its own merits.
Consequently, the only way in which these speciously-sounding challenges
can be adequately met is by removing some individuals of a species from
a state of nature, and so from all known influences of natural
selection; then, while carefully avoiding artificial selection, causing
these individuals and their progeny through many generations unduly to
exercise some parts of their bodies, or unduly to fail in the exercise
of others. But, clearly, such an experiment is one that must take years
to perform, and therefore it is now too early in the day to reproach the
followers of Darwin with not having met the challenges which are thrown
down by the followers of Weismann[31].
[31] I say "the followers of Weismann," because Weismann himself,
with his clear perception of the requirements of experimental
research, expressly states the above considerations, with the
conclusions to which they lead. Nevertheless, he is not
consistent in his utterances upon this matter; for he
frequently expresses himself to the effect, "that the _onus
probandi_ rests with my opponents, and therefore they ought to
bring forward actual proofs" (_Essays_, i. p. 390). But, as
above shown, the _onus_ rests as much with him as with his
opponents; while, even if his opponents are right, he elsewhere
recognizes that they can bring "actual proofs" of the fact only
as a result of experiments which must take many years to
perform.
Public-domain text, read in full here on John Shaqi.
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