Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
It will be at once apparent that a study of the topographical
distribution of organic types is of even more importance for us than a
study of their geographical distribution. For while the former study is
conducted, as it were, with a low power of our observing microscope, the
latter is conducted with a high power. The larger facts of geographical
distribution yield, indeed, all the general characters which we might
expect them to yield, on the theory that divergence of specific types on
common areas has been in chief part determined by physiological
conditions. But for the purpose of testing this theory in a still more
exacting manner, it is of the first importance to consider the more
detailed facts of topographical distribution, since we here come to
closer quarters with the problem of specific differentiation. Therefore,
as we have already considered this problem under the most general points
of view, we will now consider it under more special points of view.
* * * * *
It is self-evident, as we have seen in the preceding section, that the
greater the number of individuals of the same species on a given area,
the less must be the power of natural selection to split that species
into two or more allied types; because, the more crowded the population,
the greater must be the uniformitarian effect of free intercrossing.
This obvious fact has been insisted upon by several previous writers on
Darwinism; and the only reason why it has not been recognized by all
naturalists is that so few of them have observed the all-important
distinction between monotypic and polytypic evolution. The denser the
population, and therefore the greater the intercrossing and the severer
the struggle for existence within the species, the better will it be for
_transmutation_ of the species by natural selection; but the worse it
will be for _differentiation_ of the species by this form of homogamy.
On the other hand, if physiological selection be entertained as a form
of homogamy, the denser the population, the better opportunity it will
have of differentiating the species, first, because a greater number of
individuals will be present in which the physiological change may arise,
and, secondly, because, if it does arise, the severity of the struggle
for existence will _then_ give natural selection a better chance of
acting rapidly and effectually on each of the isolated sections.
Public-domain text, read in full here on John Shaqi.
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