Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
Species may be synoical at all stages of relationship. We come
across varieties, scarcely distinguishable from one another,
growing in the same locality (as, for example, the _Cirsium
heterophyllum_, with smooth or jagged leaves, the _Hieracium
sylvaticum_, with or without caulinary leaves); again, we meet
other varieties more accentuated (as the _H. hoppeanum_, with under
ligules of white or red, the _Campanula_, with white or lilac
flowers, &c.), other varieties even more marked, which might almost
be elevated to the rank of species (_Hieracium alpinum_, with hairs
and glands, and the new form _H. holadenium_, which has only
glands, _Campanula rotundifolia_ with smooth and hairy leaves), or
forms still more distinct, up to well-defined species. I could
enumerate endless examples at all stages.
It will be seen that in my definition of synoicy I do not mean to
assert that _all_ allied forms are invariably found together, but
that they are much more often seen in groups than singly. Take, for
instance, nine forms closely related (_A_ to _I_). _A_, _E_, _H_
will be found side by side at one point, _B_, _D_ at another, _C_,
_F_ at a third, &c. These facts are plainly opposed to the theory
of isolation and amixia, and make, on the contrary, in favour of
the social development of species (_loc. cit._, p. 221).
Not to multiply quotations to the same general effect, I will supply but
one other, referring to a particular case.
At one spot (_Rothwand_) much exposed to the sun, and difficult of
access, I remarked two closely allied forms, so nearly related to
_H. villosum_ that this would seem to be an intermediary form
between the two. One of these (_H. villosissimum_) is distinguished
by its tongue and thick pubescence, its tolerably large capitula,
and by the lengthened and separated scales of the involucrum; the
other, on the contrary (_H. elongatum_), is less pubescent, has
smaller capitula, and more compact scales on the involucrum than
_H. villosum_. Both are finally distinguishable from the type by
their longer stalks, which are more decidedly aphyllous, and by
their later flowering. At the spot where I found them the two forms
were closely intermingled, and each was represented by a
considerable number of plants. I did not find them anywhere else on
the mountain, nor could I find at the spot where these were growing
a single specimen of the true _H. villosum_, nor a single hybrid
from these two.
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