Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
Having devoted his life to closely observing in divers stations
multitudes of different species of plants--annuals and perennials,
bulbous and aquatic, trees and shrubs--M. Jordan has been able to
satisfy himself, and the French school of botanists to which this line
of observation has given rise, that in most cases (or "nearly
everywhere"), when a Linnean species is indigenous to a country and is
there of common occurrence, this species within that district is
represented by more or less numerous and perfectly constant varieties.
These varieties are constituted by such minute differences of
morphological character that their very existence eluded the
observation of botanists, until M. Jordan began to search specially for
them as the special objects of his scrutiny. Moreover, these varieties
of a Linnean species occupy common areas, and there grow in intimate
association with one another, or as M. Jordan says, "_pêle-mêle_." So
far, be it noticed, Jordan was proceeding on exactly the same lines as
Nägeli; only he carried his observations over a still wider range of
species on the one hand, and into a still minuter search for varieties
on the other. But the all-important point for us is, that he further
proceeded to test by experiment the physiological relations between
these morphological varieties; and found, in many hundreds of cases,
that they not only came true to seed (i. e. are hereditary and not
merely climatic), but likewise cross-sterile _inter se_. For these
reasons, M. Jordan, who is opposed to the theory of evolution, regards
all such varieties as separately created species; and the inspiring
motive of his prolonged investigations has been a desire to multiply
these proofs of creative energy. But it clearly makes no difference, so
far as evolutionists are concerned with them, whether all this multitude
of sexually isolated forms be denominated species or varieties.
The points which are of importance to evolutionists--and of the first
order of importance in the present connexion--may be briefly summarized
as follows:--
(1) The research embraces large numbers of species, belonging to very
numerous and very varied orders of plants; (2) in the majority of
cases--although not all--indigenous species which are of common
occurrence present constant varieties; (3) these varieties,
nevertheless, may be morphologically so slight as to be almost
imperceptible; (4) they occupy common areas and grow in intimate
association; (5) although many of them have undergone so small an amount
of morphological change, they have undergone a surprising amount of
physiological change; for (6) not only do very many of these varieties
come true to seed; but, (7) when they do, they are always more or less
cross-infertile _inter se_.
Public-domain text, read in full here on John Shaqi.
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