Darwiniana : Essays — Volume 02Huxley, Thomas Henry
Science
Darwiniana : Essays — Volume 02
Huxley, Thomas Henry
Darwin, Charles, 1809-1882; Evolution (Biology)
The frequent occurrence of strata containing assemblages of organic forms
which are intermediate between those of adjacent formations, is, to my
mind, fatal to this view. In the well-known St. Cassian beds, for example,
Palaeozoic and Mesozoic forms are commingled, and, between the Cretaceous
and the Eocene formations, there are similar transitional beds. On the
other hand, in the middle of the Silurian series, extensive unconformity of
the strata indicates the lapse of vast intervals of time between the
deposit of successive beds, without any corresponding change in the Fauna.
Professor Haeckel will, I fear, think me unreasonable, if I say that he
seems to be still overshadowed by geological superstitions; and that he
will have to believe in the completeness of the geological record far less
than he does at present. He assumes, for example, that there was no dry
land, nor any terrestrial life, before the end of the Silurian epoch,
simply because, up to the present time, no indications of fresh water, or
terrestrial organisms, have been found in rocks of older date. And, in
speculating upon the origin of a given group, he rarely goes further back
than the "Ante-period," which precedes that in which the remains of animals
belonging to that group are found. Thus, as fossil remains of the majority
of the groups of _Reptilia_ are first found in the Trias, they are
assumed to have originated in the "Antetriassic" period, or between the
Permian and Triassic epochs.
I confess this is wholly incredible to me. The Permian and the Triassic
deposits pass completely into one another; there is no sort of
discontinuity answering to an unrecorded "Antetrias"; and, what is more, we
have evidence of immensely extensive dry land during the formation of these
deposits. We know that the dry land of the Trias absolutely teemed with
reptiles of all groups except Pterodactyles, Snakes, and perhaps Tortoises;
there is every probability that true Birds existed, and _Mammalia_
certainly did. Of the inhabitants of the Permian dry land, on the contrary,
all that have left a record are a few lizards. Is it conceivable that these
last should really represent the whole terrestrial population of that time,
and that the development of Mammals, of Birds, and of the highest forms of
Reptiles, should have been crowded into the time during which the Permian
conditions quietly passed away, and the Triassic conditions began? Does not
any such supposition become in the highest degree improbable, when, in the
terrestrial or fresh-water Labyrinthodonts, which lived on the land of the
Carboniferous epoch, as well as on that of the Trias, we have evidence that
one form of terrestrial life persisted, throughout all these ages, with no
important modification? For my part, having regard to the small amount of
modification (except in the way of extinction) which the Crocodilian,
Lacertilian, and Chelonian _Reptilia_ have undergone, from the older
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