Disease in plantsWard, H. Marshall (Harry Marshall)
Science
Disease in plants
Ward, H. Marshall (Harry Marshall)
Plant diseases
If a suitable mass of living cells is neatly cut with a sharp razor the
first perceptible change is one of colour: the white "flesh" of a potato
or an apple, for instance, turns brown as the air enters the cut cells,
and the microscope shows that this browning affects cell-walls and
contents alike. The cut cells also die forthwith; and the oxygen of the
air combining with some of their constituents forms the brown colouring
matter which soaks into the cell-walls. The uninjured cells below them
grow longer, pushing up the dead débris, and divide across by walls
parallel to the plane of the wound, and so form series of tabular cells
with thin walls, which also soon turn brown and die, the cell-walls
meanwhile undergoing changes which convert them into cork. The living
cells deeper down are now shut off from the outer world by a skin, of
several layers, of cork-cells, which prevent the further free access of
air or moisture. During the period of active cell-division which
initiates the cork, the temperature of the growing cells rises: a sort
of fever (wound-fever) is induced, evidently owing to the active
respiration of the growing cells.
This healing by cork occurs in any tissue of living cells exposed by a
cut--leaf-tissue, young stem or root, fruit, cambium, etc.; and the same
applies to any other kind of cutting or tearing injury--such as a prick
with a needle or the proboscis of an insect, a stripping, or even a
bruise.
Such healing is prepared for and carried out very thoroughly in the case
of falling leaves and cast branches, the plane of separation being
covered by a cicatrix of cork.
If the cell-tissue under the wound is actually growing at the time,
however, a further process is observed when the wound-cork has been
formed. The uninjured cells below go on growing outwards more vigorously
than ever, the pressure of the overlying tissues taken off by the cut
having been removed, and, lifting up the cork-layer as they do so, they
rapidly divide into a juicy mass of thin-walled cells which is of a
cushion-like nature and is termed a _Callus_. This callus is at first a
homogeneous tissue of cells which are all alike capable of growing and
dividing, but in course of time it undergoes changes in different parts
which result in the formation of tracheids, vessels, fibres and other
tissue-elements, and even organs, just as the embryonic tissues of the
growing points, cambium, etc., of the healthy plant give origin to new
growths. Such wound-wood, however, is apt to differ considerably in the
arrangement, constitution and hardness of its parts as compared with
normal wood, and its peculiar density and cross-graining are often
conspicuous.
Public-domain text, read in full here on John Shaqi.
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