Dragons of the Air: An Account of Extinct Flying ReptilesSeeley, H. G. (Harry Govier)
Science
Dragons of the Air: An Account of Extinct Flying Reptiles
Seeley, H. G. (Harry Govier)
Pterosauria
The region of the back in a Pterodactyle is short as compared with the
neck, and relatively is never longer than the corresponding region in a
bird. The shortness results partly from the short length of the
vertebræ, each of which is about as long as wide. There is also a
moderate number of bones in the back. In most skeletons from Solenhofen
these vertebræ between the neck and girdle of hip bones number from
twelve to sixteen. They have a general resemblance in form to the dorsal
vertebræ in birds. The greatest number of such vertebræ in birds is
eleven. The number is small because some of the later vertebræ in birds
are overlapped by the bones of the hip girdle, which extend forward and
cover them at the sides, so that they become blended with the sacrum.
This region of the skeleton in the Dimorphodon from the Lias is
remarkable for the length of the median process, named the neural spine,
which is prolonged upward like the spines of the early dorsal vertebræ
of Horses, Deer, and other mammals. In this character they differ from
living reptiles, and parallel some Dinosaurs from the Weald. The bones
of the back in Ornithocheirus from the Cambridge Greensand show the
under side to be well rounded, so that the articular surfaces between
the vertebræ, though still rather wider than deep, are much less
depressed than in the region of the neck. The neural canal for the
spinal cord has become larger and higher, and the sides of the bone are
somewhat compressed. Strong transverse processes for the support of the
ribs are elevated above the level of the neural canal, at the sides of
vertebræ compressed on the under sides, and directed outward. Between
these lateral horizontal platforms is the compressed median neural
spine, which varies in vertical height. The articulation of the ribs is
not seen clearly. Isolated ribs from the Stonesfield Slate have
double-headed dorsal ribs, like those of birds. In some specimens from
the Solenhofen Slate like the Scaphognathus, in the University Museum at
Bonn, dorsal ribs appear to be attached by a notch in the transverse
process of the dorsal vertebra, which resembles the condition in
Crocodiles. Variations in the mode of attachment of ribs among mammals
may show that character to be of subordinate importance. Von Meyer has
described the first pair of ribs as frequently larger than the others,
and there appear in Rhamphorhynchus to be examples preserved of the
sternal ribs, which connect the dorsal ribs with the sternum. Six pairs
have been counted. A more interesting feature in the ribs consists in
the presence behind the sternum, which is shorter than the corresponding
bone in most birds, of median sternal ribs. They are slender #V#-shaped
bones in the middle line of the abdomen, which overlapped the ends of
the dorsal ribs like the similar sternal bones of reptiles. Such
structures are unknown among Birds and Mammals. There is no trace in the
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