Dragons of the Air: An Account of Extinct Flying ReptilesSeeley, H. G. (Harry Govier)
Science
Dragons of the Air: An Account of Extinct Flying Reptiles
Seeley, H. G. (Harry Govier)
Pterosauria
On the left-hand side the two prepubic bones are separate. On the
right-hand they are united into a transverse bar which overlaps the
front of pelvis seen from the under side]
There are three modifications of the Ornithosaurian pelvis. First, the
type of Rhamphorhynchus, in which the pubis and ischium are inclined
somewhat backward, and in which the two prepubic bones are triangular,
and are often united together to form a transverse bow in front of the
pubic region.
Secondly, there is the ordinary form of pelvis in which the pubis and
ischium usually unite with each other down their length, as in
Dimorphodon, but sometimes, as in _Pterodactylus dubius_, divide
immediately below the acetabulum. All these types possess the
paddle-shaped prepubic bones, which are never united in the median line.
Thirdly, there is the cretaceous form indicated by Ornithocheirus and
Ornithostoma, in which the posterior half of the ilium is modified in a
singular way, since it is more elevated towards the sacrum than the
anterior half, suggesting the contour of the upper border of the ilium
in a lizard. Without being reptilian--the anterior prolongation of the
bone makes that impossible--it suggests the lizards. This type also
possesses prepubic bones. They appear, according to Professor Williston,
to be more like the paddle-shaped bones of Pterodactylus than like the
angular bones in Rhamphorhynchus. The prepubic bones are united in the
median line as in Rhamphorhynchus. But their median union in that genus
favours the conclusion that the bones were united in the median line in
all species, though they are only co-ossified in these two families.
[Illustration: FIG. 32. THE PELVIC BONES OF AN ALLIGATOR SEEN FROM
BELOW
The bones in front are here regarded as prepubic, but are commonly
named pubic]
This median union of the prepubic bones is a difference from those
mammals like the Ornithorhynchus and Echidna, which approach nearest to
the Reptilia. In them the prepubic bones have a long attachment to the
front margin of the pubis, and extend their points forward without any
tendency for the anterior extremities to approximate or unite. The
marsupial mammals have the same character, keeping the marsupial bones
completely distinct from each other at their free extremities. The only
existing animals in which an approximation is found to the prepubic
bones in Pterodactyles are Crocodiles, in bones which most writers term
the pubic bones. This resemblance, without showing any strong affinity
with the Crocodilia, indicates that Crocodiles have more in common with
the fossil flying animals than any other group of existing reptiles; for
other reptiles all want prepubic bones, or bones in front of the pubic
region.
THE HIND LIMB
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