Earthworms and Their AlliesBeddard, Frank E. (Frank Evers)
Science
Earthworms and Their Allies
Beddard, Frank E. (Frank Evers)
Earthworms
Oceanic islands are islands that have always been islands, a
definition that seems tautological until we compare it with some
other land masses that may be termed 'islands.' Geology teaches us
in fact that from the point of view of their origin islands may be
divided into two quite sharply contrasted classes. There are those
detached land masses usually lying near to or comparatively near to
some continent, which have been in the course of time detached by the
action of the waves from that continent, such as for instance the
British Isles, which undoubtedly represent a portion of the European
continent which was once quite continuous with Europe. On the other
hand we have the Hawaiian archipelago, St Helena, Fernando Noronha,
and other similar islands, which are more remote in their position
from continents and concerning which it seems clear that they have
originated _de novo_ by the action of submarine volcanos or of the
growth of coral, combined with subsidence, following elevation, or
from several of the causes combined. In any case the islands which
are termed oceanic islands have never formed part of a continent.
They are not relics of previously existing continents. It becomes
a matter of great interest to compare the earthworms which are to
be found upon oceanic islands with those which inhabit continental
islands. Fortunately there are a good many facts at our disposal for
this purpose; and we shall compare the earthworms of the Hawaiian
archipelago with those which are found upon certain small islands
lying to the south of New Zealand, viz. Campbell and Auckland islands
and the more southern Macquarie islands.
The earthworms of the Hawaiian archipelago have been studied by a
good many persons, and altogether a number of species have been
described from that group of islands of which the following is a
list: _Pheretima hawayana_, _P. heterochaeta_, _P. peregrina_,
_P. schmardae_, _P. hesperidum_, _P. morrisi_, _P. perkinsi_,
_P. biserialis_ (= _P. elongata_), _Allolobophora putris_ (=
Kinberg's _Hypogaeon havaicum_), _A. foetida_, _A. caliginosa_, _A.
nordenskiöldi_, _A. limicola_, _A. rosea_, and finally the well-known
_Pontoscolex corethrurus_. Of these species there is only one which
is even possibly a form limited to the Sandwich Islands, and that is
_Pheretima perkinsi_, a species which I myself at first described as
a new form, but which was afterwards regarded as identical with _P.
heterochaeta_ by Michaelsen, and later still resuscitated by Ude. All
the others are found in many parts of the world and not only in the
nearest mainland to the archipelago which we are now considering.
I have had already occasion to speak of some of them as peregrine
forms, especially of _Pontoscolex corethrurus_ which occurs all over
the world.
Public-domain text, read in full here on John Shaqi.
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