=151. Influence of light on the movement of chlorophyll
bodies.=—_In fern prothallia_.—If we place fern prothallia in weak
light for a few hours, and then examine them under the microscope,
we find that the most of the chlorophyll bodies in the cells are
arranged along the inner surface of the horizontal wall. If now the
same prothallia are placed in a brightly lighted place for a short
time most of the chlorophyll bodies move so that they are arranged
along the surfaces of the perpendicular walls, and instead of having
the flattened surfaces exposed to the light as in the former case, the
edges of the chlorophyll bodies are now turned toward the light. (See
figs. 70, 71.) The same phenomenon has been observed in many plants.
Light then has an influence on chlorophyll bodies, to some extent
determining their position. In weak light they are arranged so that the
flattened surfaces are exposed to the incidence of the rays of light,
so that the chlorophyll will absorb as great an amount as possible
of kinetic energy; but intense light is stronger than necessary, and
the chlorophyll bodies move so that their edges are exposed to the
incidence of the rays. This movement of the chlorophyll bodies is
different from that which takes place in some water plants like elodea.
The chlorophyll bodies in elodea are free in the protoplasm. The
protoplasm in the cells of elodea streams around the inside of the cell
wall much as it does in nitella and the chlorophyll bodies are carried
along in the currents, while in nitella they are stationary.
CHAPTER VIII.
STARCH AND SUGAR CONCLUDED. ANALYSIS OF PLANT SUBSTANCE.
1. Translocation of Starch.
=152. Translocation of starch.=—It has been found that leaves of
many plants grown in the sunlight contain starch when examined after
being in the sunlight for several hours. But when the plants are left
in the dark for a day or two the leaves contain no starch, or a much
smaller amount. This suggests that starch after it has been formed may
be transferred from the leaves, or from those areas of the leaves where
it has been formed.
[Illustration: Fig. 72.
Leaf of tropæolum with portion covered with corks to prevent the
formation of starch. (After Detmer.)]
[Illustration: Fig. 73.
Leaf of tropæolum treated with iodine after removal of cork, to show
that starch is removed from the leaf during the night.]
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