When the cells have absorbed a certain amount of a given substance,
no more is absorbed until the concentration of the cell-sap in that
particular substance is reduced. This, however, does not interfere
with the absorption of water, or of other substances in solution by
the same cells. Plants have therefore a certain selective power in the
absorption of food substances.
=174. Action of root hairs on insoluble substances. Acidity of root
hairs.=—If we take a seedling which has been grown in a germinator,
or in the folds of cloths or paper, so that the roots are free from the
soil, and touch the moist root hairs to blue litmus paper, the paper
becomes red in color where the root hairs have come in contact. This
is the reaction for the presence of an acid salt, and indicates that
the root hairs excrete certain acid substances. This acid property of
the root hairs serves a very important function in the preparation
of certain of the elements of plant food in the soil. Certain of the
chemical compounds of potash, phosphoric acid, etc., become deposited
on the soil particles, and are not soluble in water. The acid of the
root hairs dissolves some of these compounds where the particles of
soil are in close contact with them, and the solutions can then be
taken up by the roots. Carbonic acid and other acids are also formed in
the soil, and aid in bringing these substances into solution.
=175.= This corrosive action of the roots can be shown by the
well-known experiment of growing a plant on a marble plate which is
covered by soil. In lieu of the marble plate, the peas may be planted
in clam or oyster shells, which are then buried in the soil of the
pot, so that the roots of the seedlings will come in contact with the
smooth surface of the shell. After a few weeks, if the soil be washed
from the marble where the roots have been in close contact, there will
be an outline of this part of the root system. Several different acid
substances are excreted from the roots of plants which have been found
to redden blue litmus paper by contact. Experiments by Czapek show,
however, that the carbonic acid excreted by the roots has the power of
directly bringing about these corrosion phenomena. The acid salts are
the substances which are most actively concerned in reddening the blue
litmus paper. They do not directly aid in the corrosion phenomena. In
the soil, however, where these compounds of potash, phosphoric acid,
etc., are which are not soluble in water, the acid salt (primary acid
potassium phosphate) which is most actively concerned in reddening
the blue litmus paper may act indirectly on these mineral substances,
making them available for plant food. This salt soon unites with
certain chlorides in the soil, making among other things small
quantities of hydrochloric acid.
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