The haustoria of the dodder form a complete junction with the vascular
bundles of its host so that through the vessels water and salts are
obtained, while through the junction of sieve tubes the elaborated
organic food is obtained. The union of the dodder with its host is like
that between a graft and the graft stock. The beech drops (Epiphegus)
is another example of a parasitic seed plant. It is parasitic on the
roots of the beech.
[Illustration: Fig. 74. Dodder.]
=180. The mistletoe= (Viscum album), which grows on the branches
of trees, sends its roots into the branches, and only the vessels
of the vascular system are fused according to some. If this is true
then it probably obtains only water and salts from its host. But the
mistletoe has green leaves and is thus able to assimilate carbon
dioxide and manufacture its own organic substances. It is claimed by
some, however, that the host derives some food from the parasite during
the winter when the host has shed its leaves, and if this is true it
would seem that organic food could also be derived during the summer
from the host by the mistletoe.
=181. Saprophytes.=—A saprophyte is a plant which is enabled
to obtain its food, especially its organic food, directly from dead
animals or plants or from dead organic substances. Many fungi are
saprophytes, as the moulds, mushrooms, etc. (See Nutrition of the
Fungi.)
=182. Humus saprophytes.=—The action of fungi as described in the
preceding chapter, as well as of certain bacteria, gradually converts
the dead plants or plant parts into the finely powdered brown substance
known as _humus_. In general the green plants cannot absorb organic
food from humus directly. But plants which are devoid of chlorophyll
can live saprophytically on this humus. They are known as _humus
saprophytes_. Many of the mushrooms and other fungi, as well as some
seed plants which lack chlorophyll or possess only a small quantity,
are able to absorb all their organic food from humus. It is uncertain
whether any seed plants can obtain all of their organic food directly
from humus, though it is believed that many can so obtain a portion
of it. But a number of seed plants, like the Indian-pipe (Monotropa)
and certain orchids, obtain organic food from humus. These plants lack
chlorophyll and cannot therefore manufacture their own carbohydrate
food. Not being parasitic on plants which can, as in the case of the
dodder and beech drops mentioned above, they undoubtedly derive their
organic food from the humus. But fungus mycelium growing in the humus
is attached to their roots, and in some orchids enters the roots and
forms a nutritive connection. The fungus mycelium can absorb organic
food from the humus and in some cases at least can transfer it over to
the roots of the higher plant (see Mycorhiza).
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