=194. Endotrophic mycorhiza.=—These are found on many of the
humus saprophytes, which are devoid of chlorophyll, as well as on those
possessing little or even on some plants possessing an abundance, of
chlorophyll. Examples are found in many orchids (see the coral root
orchid, for example), some of the ferns (Botrychium), the pines,
leguminous plants, etc. In endotrophic mycorhiza the mycelium is more
abundant within the tissues of the root, though some of the threads
extend to the outside. In the case of the mycorhiza on the humus
saprophytes which have no chlorophyll, or but little, it is thought
by some that the fungus mycelium in the humus assists in converting
organic substances and carbohydrates into a form available for food
by the higher plant and then conducts it into the root, thus aiding
also in the process of absorption, since there are few or no root
hairs on the short and fleshy mycorhiza. The roots, however, of some
of these humus saprophytes have the power of absorbing a portion of
their organic compounds from the humus. It is thought by some, though
not definitely demonstrated, that in the case of the oaks, beeches,
hornbeans, and other chlorophyll-bearing symbionts, the fungus threads
do not absorb any carbohydrates for the higher symbiont, but that they
actually derive their carbohydrates from it.[14] But it is reasonably
certain that the fungus threads do assimilate from the humus certain
unoxidized, or feebly oxidized, nitrogenous substances (ammonia, for
example), and transfer them over to the host, for the higher plants
with difficulty absorb these substances, while they readily absorb
nitrates which are not abundant in humus. This is especially important
in the forest. It is likely therefore.
5. Nitrogen gatherers.
[Illustration: Fig. 81. Root of the common vetch, showing root
tubercles.]
=195. How clovers, peas, and other legumes gather nitrogen.=—It
has long been known that clover plants, peas, beans, and many other
leguminous plants are often able to thrive in soil where the cereals
do but poorly. Soil poor in nitrogenous plant food becomes richer in
this substance where clovers, peas, etc., are grown, and they are often
planted for the purpose of enriching the soil. Leguminous plants,
especially in poor soil, are almost certain to have enlargements, in
the form of nodules, or “root-tubercles.” A root of the common vetch
with some of these root-tubercles is shown in fig. 81.
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