Elements of Structural and Systematic Botany: For High Schools and Elementary College CoursesCampbell, Douglas Houghton
Science
Elements of Structural and Systematic Botany: For High Schools and Elementary College Courses
Campbell, Douglas Houghton
Botany -- Textbooks
The spores arise from a single layer of cells near the outside of
the inner mass of cells (_G_, _sp._). These cells (_H_, _sp._) are
filled with glistening, granular protoplasm; have a large and
distinct nucleus, and no chlorophyll. They finally become entirely
separated and each one gives rise to four spores which closely
resemble those of the liverworts but are smaller.
Near the base of the capsule, on the outside, are formed breathing
pores (Fig. 58, _F_) quite similar to those of the higher plants.
If the spores are kept in water for a few days they will germinate,
bursting the outer brown coat, and the contents protruding through
the opening surrounded by the colorless inner spore membrane. The
protuberance grows rapidly in length and soon becomes separated from
the body of the spore by a wall, and lengthening, more and more,
gives rise to a green filament like those we found attached to the
base of the full-grown plant, and like those giving rise to buds
that develop into leafy plants.
CLASSIFICATION OF THE MOSSES.
The mosses may be divided into four orders: I. The peat mosses
(_Sphagnaceæ_); II. _Andreæaceæ_; III. _Phascaceæ_; IV. The common
mosses (_Bryaceæ_).
[Illustration: FIG. 64.--_A_, a peat moss (_Sphagnum_), × ½. _B_, a
sporogonium of the same, × 3. _C_, a portion of a leaf, × 150. The
narrow, chlorophyll-bearing cells form meshes, enclosing the large,
colorless empty cells, whose walls are marked with thickened bars, and
contain round openings (_o_).]
The peat mosses (Fig. 64) are large pale-green mosses, growing often
in enormous masses, forming the foundation of peat-bogs. They are of a
peculiar spongy texture, very light when dry, and capable of absorbing
a great amount of water. They branch (Fig. 64, _A_), the branches
being closely crowded at the top, where the stems continue to grow,
dying away below.
[Illustration: FIG. 65.--Forms of mosses. _A_, plant of _Phascum_,
× 3. _B_, fruiting plant of _Atrichum_, × 2. _C_, young capsule of
hairy-cap moss (_Polytrichum_), covered by the large, hairy calyptra.
_D_, capsules of _Bartramia_: i, with; ii, without the calyptra. _E_,
upper part of a male plant of _Atrichum_, showing the flower, × 2.
_F_, a male plant of _Mnium_, × 4. _G_, pine-tree moss (_Clemacium_),
× 1. _H_, _Hypnum_, × 1. _I_, ripe capsules of hairy-cap moss: i,
with; ii, without calyptra.]
The sexual organs are rarely met with, but should be looked for late
in autumn or early spring. The antheridial branches are often
bright-colored, red or yellow, so as to be very conspicuous. The
capsules, which are not often found, are larger than in most of the
common mosses, and quite destitute of a stalk, the apparent stalk
being a prolongation of the axis of the plant in the top of which the
base of the sporogonium is imbedded. The capsule is nearly globular,
opening by a lid at the top (Fig. 64, _B_).
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